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Fulmars (Fulmarus glacialis) on water, feeding at a fish processing plant outfall. Grundafjordur, Iceland, March
Ptarmigan (Lagopus mutus) in winter plumage in flight, Utsjoki, Finland, April
Sycamore tree (Acer pseudoplatanus), growing on limestone pavement, Lancashire, UK. May
Rock formations silhouetted at sunset on the Pacfic coast of Olympic National Park, Washington State, USA
Icicles hanging from sandstone cliffs on shoreline, Apostle Islands National Lakeshore, Lake Superior, Squaw Bay, Wisconsin, February 2014
Water rushing down alpine stream, Logan Pass, Glacier National Park, Rocky Mountains, Montana, July 2010
Giants Causeway, looking in to land, Causeway coast, Antrim county, Northern Ireland, UK, September 2013
Fissure eruption of Chico Volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Vancouver island black bears (Ursus americanus vancouveri) taken with remote camera, Vancouver Island, British Columbia, Canada, August
Sandstone cliff at sunset, Colorado Plateau, Kodachrome Basin State Park, Utah, USA November 2012
Sandstone formations in Goblin Valley State Park, Utah, USA November 2012
Rufous elephant shrews (Elephantulus rufescens) Botswana
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, walking down a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Red / Blotcheye soldierfish (Myripristis murdjan) and sea whips coral, Junceella at Fotteyo caves, Maldives, Indian Ocean
Harlequin duck (Histrionicus histrionicus) male on a large boulder in the fast flowing and turbulent waters of the Laxa River, Iceland, May. Bookplate from Danny Greens The Long Journey North
Atlantic puffins (Fratercula arctica) at dusk, Fair Isle, Shetland, July. Bookplate from Danny Greens The Long Journey North
Puffin (Fratercula arctica) stretching wings, late evening light. Shetland Islands, Scotland, July
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January
Bengal Tigers (Panthera tigris tigris) resting in the shade national park, India, taken on location for Tiger - Spy in the Jungle March 2007
Old Harry Rocks at dawn, looking towards the Isle of Wight, Studland, Dorset, England, UK. September 2015
RF - Old Harry Rocks, colourful sunrise looking towards the Isle of Wight, Studland, Dorset, UK. September
Trevose Lighthouse at dusk, long exposure at high tide, Trevose, Cornwall, UK. January 2015
RF- Grey Wolf (Canis lupus) howling at sunset (digitally enhanced)
Ice splash out formation with icicles, Lake Baikal, Siberia, Russia, March
Antelope Canyon, a slot canyon with eroded sandstone patterns, Navajo Tribal Park, Arizona, USA, December 2012
Black-backed Jackal (Canis mesomelas) drinking, Kgalagadi Transfrontier Park, South Africa. January
Sandstone monoliths, Colorado Plateau, Kodachrome Basin State Park, Utah, USA November 2012
Snowy owl (Bubo scandiaca) in landscape, Utsjoki, Finland, April
Common hamster feeding (Cricetus cricetus) Alsace, France, April, captive
Gannet (Morus bassanus) adult diving, Shetland Islands, Scotland, UK, July
Cliff top view of a gannet (Morus bassanus) colony. Shetland Islands, Scotland, UK, July
Cracked mud and blue stones, Fuerteventura, Canary Islands
Stone on rock pattern, Kilve beach, Somerset, UK. January 2014
Waikoropupu Springs, Golden Bay, South Island, New Zealand
Bamburgh Castle, early morning light at dawn, Bamburgh, Northumberland, UK, April 2013
Eroded sandstone patterns on walls of Lower Canyon, Antelope Canyon, Page, Arizona, USA, February 2015
Marine Iguana (Amblyrhynchus cristatus) on rocks with wave splash. Galapagos, Ecuador
View looking up a waterfall from within a stream, Snowdonia NP, Gwynedd Wales, UK, December
A split-level image of a mountain stream, in autumn, with water droplets on lens, Nant Ffrancon Valley, Snowdonia NP, Gwynedd, Wales, UK, October
A split-level view through a small waterfall, Llanberis, Snowdonia NP, Gwynedd, Wales, UK, July
Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
A view of a forested inlet lit by low sunlight. Aggro Beach, west coast of Vancouver Island, Canada, September 2010
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)
Harlequin duck (Histrionicus histrionicus) portrait of male stood motionless on exposed rock, in river, Iceland, May
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
California sea lion pup (Zalophus californianus) portrait in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Dipper (Cinclus cinclus) landing on exposed stone, with insect prey for chicks in beak, Brecon Beacons National Park, Wales, UK
Map lichen (Rhizocarpon geographicum) on rock with (Hyoseris radiata) flower, Menorca, Balearic Islands, Spain, Europe