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RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
RF - Reindeer (Rangifer tarandus) on a snowy ridge, Svalbard, Norway, April
RF- Great grey owl (Strix nebulosa) in flight, Finland, March
RF- Razorbill (Alca torda) resting on rocks. Great Saltee Island, County Wexford, Ireland. April
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
RF- Eurasian Eagle owl (Bubo bubo) adult perched on rocky outcrop with the Super Full Moon, September 28th 2015, Southern Norway. August. Multiple exposure
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
RF - Red Squirrel (Sciurus vulgaris) adult in summer coat leaping onto fallen log. Scotland, UK. September
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
RF- Serotine bat (Vespertilio serotinus) in flight above Hogweed. Captive, UK
RF- Red squirrel in autumn (Sciurus vulgaris). Strathspey, Scotland, UK
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF - Green sea turtle (Chelonia mydas) beneath clouds. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
RF- Peregrine Falcon (Falco peregrinus) landing on moorland with its wings spread. Scotland, UK, March
RF - Early Purple Orchid (Orchis mascula) in flower, Broxwater, Cornwall, UK. May 2016
RF- Sunset on Kimmeridge Bay, Isle of Purbeck, Jurassic Coast World Heritage Site, Dorset, England, UK. November 2008
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
RF - Marbled White butterfly (Melanargia galathea) resting on common spotted orchid, Dunsdon Nature Reserve, Devon, UK. July
RF- Daffodils (Narcissus) in flower photographed using soft focus technique
RF- Pool Frog (Rana lessonae) sitting on white lily pad, Danube delta rewilding area, Romania
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
RF- Small jellyfish (Pelagia noctiluca) swimming beneath surface. Marine Protected Area of Portofino (Area Marina Protetta, Portofino), Liguria, Italy. Mediterranean Sea
RF - Long-spined sea urchin (Astropyga radiata). Lembeh Strait, North Sulawesi, Indonesia
RF- Close up of Leaves of Red Oak (Quercus rubra) in autumn. October
RF- Hyacinth macaw (Anodorhynchus hyacinthinus) in its nest hole. Pousada Aguape Lodge, Mato Grosso do Sul State, Brazil. September
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
RF - Granular poison frog (Oophaga granulifera). Osa Peninsula, Costa Rica. Vulnerable species
RF - Lowland rainforest, Osa Peninsula, Costa Rica
RF - Harts tongue fern (Phyllitis scolopendrium) leaf unfurling and creating the shape of a heart, Cornwall, UK, May
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
RF - Blue waterlily (Nymphaea caerulea) close up into centre of flower. Occurs in Africa and Asia. Botanic Garden Meise, Belgium, August
RF - Close up of Greater stitchwort (Stellaria holostea) flower, Cornwall, UK. April
RF- Grape hyacinths (Muscari) and Daffodils (Narcissus sp) in flower, Norfolk, UK, March
RF- Grey seal (Halichoerus grypus) pups on Norfolk Beach, England, UK, December
RF- European hedgehog (Erinaceus europaeus) orphan, Jarfalla, Sweden. August
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
RF- Siberian tiger (Panthera tigris altaica) young cub, lying down, looking away, captive
RF- Tiger (Panthera tigris) 14-month Lakshmi cub resting in tree, Bandhavgarh National Park, India. Endangered species
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species