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Rhododendron (Rhododendron sp) in flower, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Temmincks tragopan (Tragopan temminckii) male bird walking throught the forest of Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Rhododendron (Rhododendrons sp) flowering, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Tibetan macaque (Macaca thibetana) female with baby, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Small waterfalls, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Golden pheasant (Chrysolophus pictus) male jumping, Yangxian Biosphere Reserve, Shaanxi, China
White eared pheasant (Crossoptilon crossoptilon) spreading his wings while jumping down a rock, Baima Snow Mountain Nature reserve, Yunnan, China, October
Red-billed blue magpie (Urocissa erythroryncha) perched on a branch, Yangxian Biosphere Reserve, Shaanxi, China, April
Wreathed hornbill (Aceros undulatus) taking off, Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China. April
Oriental Pied hornbill (Anthracoceros albirostris) male bringing berry to nest, Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Grey peacock-pheasant (Polyplectron bicalcaratum) close up of the feathers, Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Two Blood pheasant birds (Ithaginis cruentus) males Baima Snow Mountain Nature Reserve, Yunnan, China. October
Cloud forest of the Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Black capped chickadee (Poecile atricapillus) portrait, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Puma (Puma concolor) resting, Corcovado National Park, Costa Rica, May
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Bairds tapir (Tapirus bairdii) walking along beach in Corcovado National Park, Costa Rica. Endangered
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Jaguar (Panthera onca) camera trap image, Tortuguero National Park, Costa Rica
Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica
Eyespots on wing of Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica, February 2015
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Sedge Warbler (Acrocephalus schoenobaenus) Greylake RSPB Reserve, Somerset Levels, England, UK, August
Guanaco (Lama guanicoe) with lake and mountains, Torres del Paine National Park, Patagonia. Chile
Domesticated Alpaca / Vicugna (Lama / Vicungna pacos) on plains with snow-capped peak in distance. Sajama National Park, Bolivia
Ascidian (Rhopalaea sp) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cheetah male (Acinonyx jubatus) in morning light. Moremi National Park, Okavango delta, Botswana, Southern Africa
Mining bee (Andrena sp) flying to Giant harebell (Campanula latifolia), Pentwyn farm SSSI, Gwent Wildlife Trust Reserve, Monmouthshire Wales UK, July
Two Cheetah (Acinonyx jubatus) chasing Springbok (Antidorcas marsupialis) Kgalagadi, South Africa
Cheetah (Acinonyx jubatus) mother and cubs 6 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) brothers looking in opposite directions, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) cubs grooming each other, Masai-Mara Game Reserve, Kenya. Vulnerable species
Underwater view of African tigerfish (Hydrocynus vittatus), predatory fish in the Okavango river with Water lily (Nymphaea nouchali var. caerulea) and other aquatic plants
Underwater view of Sungai Melinau River, Gunung Mulu National Park, Sarawak, Borneo, Malaysia. Photographed for The Freshwater Project
Cheetah (Acinonyx jubatus) cubs aged 9 months playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) mother and cub aged 9 months on termite mound, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) cubs aged 5 weeks, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) cub sitting portrait, Serengeti National Park, Tanzania
Rangimairewhenua or Blue Lake, sacred to the Maori, Nelson Lakes National Park, South Island, New Zealand, February 2013. Photographed for The Freshwater Project
White-rumped Snowfinch (Montifringilla taczanowskii) with chick at nest burrow, Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau
Silver Birch (Betula pendula) in autumn, Craigellachie National Nature Reserve, Speyside, Cairngorms National Park, Scotland, October
Reindeer (Rangifer tarandus) bull reindeer with antlers in velvet, reintroduced Cairngorm Reindeer Herd, Cairngorm National Park, Speyside, Scotland, October
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
RF - Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
Hanuman langur (Semnopithecus / Presbytis entellus), female with baby at water tapHanuman langur (Semnopithecus/ Presbytis entellus), female with baby at water tap, Ranthambhore National Park, Rajasthan, India
Smooth-coated otter (Lutrogalle perspicillate) about to enter a marsh at dawn, Dudhwa National Park, Uttar Pradesh, India. Photographed using a camera trap