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Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) amongst seaweed on a maerl bed, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
View towards Higger Tor from Over Owler Tor with heather in full bloom. Abandoned millstone can be seen in the foreground. Peak District National Park, Derbyshire, UK. August 2015
Brimstone butterfly (Gonepteryx rhamni) on Devil s-bit Scabious (Succisa pratensis). Peak District National Park, Derbyshire, UK. August
Harebells (Campanula rotundifolia) growing in disused limestone quarry. Peak District National Park, Derbyshire, UK. August
The Pinnacle Stone, Curbar Edge surrounded by flowering heather. Peak District National Park, Derbyshire, UK. August
Male New Forest Pony looking alert and listening with ears pricked standing among blooming Common Heather or Ling (Calluna vulgaris) on Hampton Ridge, New Forest National Park, Hampshire England, UK
Sika deer (Cervus nippon) stags in summer coat, Arne RSPB Reserve, Dorset, England, August
Juvenile European beaver {Castor fiber} striping bark off a Birch trunk at the Aigas Field Studies Centre, European Beaver demonstration project, Inverness-shire, Scotland, August 2008
High altitude Boeri Lake in old volcano crater, Morne Trois Pitons National Park, Dominica, West Indies, Windward Islands
Old trees in the Syndicate Forest National Park, Dominica, Eastern Caribbean. February 2017
Alpine Ibex (Capra ibex) in its landscape, Valsavarenche, Gran Paradiso national park, Aosta Valley, Italy, May
Red fox (Vulpes vulpes) in deep snow emerging from den, Valsavarenche valley, Gran Paradiso national Park, Aosta Valley, Alps, Italy, December
Alpine marmot (Marmota marmota) carrying grass and other nesting material across snow to its nest while another one is on the lookout, Gran Paradiso National Park, Aosta Valley, Italy, April
Delacouras langurs (Trachypithecus delacouri) restin, Van Long Nature Reserve, Ninh Binh, VietnamDelacours langurs (Trachypithecus delacouri) restin, Van Long Nature Reserve, Ninh Binh, Vietnam. Critically endangered
Red shanked douc langur (Pygathrix nemaeus) Son Tra Nature Reserve, Vietnam
Chimpanzee female (Pan troglodytes schweinfurthii) sleeping in a nest built in a tree, Kibale National Park, Uganda, January
Female Uganda kob (Kobus kob thomasi) suckling calf. Murchisson Falls National Park, Uganda. February
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Coalition of two African male lions (Panthera leo) resting on the banks of the Luangwa river, dry season, South Luangwa National Park, Zambia
Hawthorn tree and granite outcrop at Holwell tor, sunrise and mist, Dartmoor National Park, Devon, UK. October 2018
Marsh fritillary butterfly (Euphydrayas aurinia) feeding on Meadow thistle flower (Cirsium helenoides), Dunsdon Nature Reserve, near Holsworthy, Devon, UK. June 18
Great Mis Tor covered in snow, Dartmoor National Park, Devon, England, UK, January 2015
Great Staple Tor at dusk, Dartmoor National Park, Devon, England, UK, January
Landscape view of Great Staple Tor, Dartmoor National Park, Devon, England, UK, January 2012
Moorland view from Littaford Tor, with the enlarged shadow of the photographer, also known as a Brocken Spectre, with a Glory caused by refracting water droplets cast onto mist by the rising sun
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male perched in marsh vegetation (Phragmites) in foreground. Bear River Migratory Bird Refuge, view toward east
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Short-eared Owl (Asio flammeus) shaking after preening while perched on fencepost, Boxelder County, Utah, USA, May
Painted bunting (Passerina ciris) male singing, Wichita Mountains National Wildlife Refuge, Oklahoma, USA, May
Brown Pelican (Pelecanus occidentalis) portrait, Pacific race, breeding plumage, Bolsa Chica Ecological Reserve, California, USA
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) two at waters edge, one shaking off water, Kiskunsagi National Park, Pusztaszer, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Forested island on Loch Lomond, Loch Lomond and the Trossachs National Park, Scotland, UK
Large-eared pika (Ochotona macrotis) Serxu County, Garze Prefecture, Sichuan Province, China
White-lipped deer (Cervus albirostris) stags in mountain landscape, Serxu County, Garze Prefecture, Sichuan Province, China
Temmincks tragopan (Tragopan temminckii) Tangjiahe Nature Reserve, Sichuan province, China
White-lipped deer (Cervus albirostris) male, Serxu, Garze Prefecture, Sichuan Province, China
Injured Muntjac (Muntiacus reevesi) swimming in a small creek after unsuccessfully attempting to jump across it. This muntjac was unable to get out of the creek himself
Pere Davids deer / Milu (Elaphurus davidianus) during the rutting season, stag, Hubei Tian ezhou Milu National Nature Reserve, Shishou, Hubei, China
Snow Leopard (Panthera uncia) female with two cubs climbing a canyon cliff at dusk at 4400 metres, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Mountain Gorilla (Goriila beringei) resting, Volcanoes National Park, Virunga Mountains, Rwanda
Cuban emerald hummingbird (Chlorostilbon ricordii) Guanahacabibes Peninsula National Park, Cuba
Bee hummingbird (Mellisuga helenae) feeding from flower, Guanahacabibes Peninsula National Park, Cuba
Orang utan (Pongo pygmaeus) baby climbing in trees, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered. June
Orang utan (Pongo pygmaeus) baby sleeping in the arms of an adult, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered