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Red grouse (Lagopus lagopus) on moorland at sunrise, Peak District National Park, UK. August
Two Rhinoceros hornbills (Buceros rhinoceros) perched in tree canopy looking at each other, Halabala Wildlife Sanctuary, Narathiwat Province, Thailand
Northern parula (Parula americana) male singing from flowering crabapple (Malus sp)Northern parula (Parula americana) male singing from flowering crabapple (Malus sp.) tree in spring, near Salamanca, New York, USA, May 2022
Jaguar (Panthera onca) walking through a lagoon in the mangroves, Marismas Nacionales, Nayarit, Mexico
RF - Cheetah (Acinonyx jubatus) female patrolling territory. Long Gully, Ngorongoro Conservation Area, Tanzania. April
Marsh deer (Blastocerus dichotomus) walking through vegetation, Pocone, Brazil
Spinifex pigeon (Geophaps plumifera) flock on stony desert floor, Diamantina River, Queensland, Australia
Group of Grey-headed flying fox (Pteropus poliocephalus) roosting, Toowoomba, Queensland, Australia
Ocelot (Leopardus pardalis) sitting on forest floor, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Roe deer (Capreolus capreolus) doe, leaping whilst running away, rear view, Vosges, France
Mallard (Anas platyrhynchos) shaking its wings with slow shutter speed to reveal movement, Martin Mere WWT Reserve, Lancashire, UK
Strinatii lungless salamander (Speleomantes strinatii) hiding under chestnut leaves into a deciduous wood in Northern Italy. San Bartolomeo di Savignone, Italy. October
RF - Red fox (Vulpes vulpes) in deep snow. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
Southern red-billed hornbill male (Tockus rufirostris) Mahatsu Reserve, Botswana, June
King penguin (Aptenodytes patagonicus) colony, Volunteer Point, East Falkland, Falkland Islands, October
Green Turtle (Chelonia mydas) returning to the sea, IUCN Endangered, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa
Short-eared owl (Asio flammeus) perched, Donana Natural Park, Andalusia, Spain. January
Corn bunting (Miliaria calandra) perched in brassica flowers, Sierra de Grazalema Natural Park, southern Spain, April
European greenfinch (Carduelis chloris), Andalusia, Spain, June
African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Collared treepie (Dendrositta frontalis) bird perched on a tree Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Hippopotamus (Hippopotamus amphibius) male marking territory with faeces, Masai Mara Game Reserve, Kenya
Crested drongo (Dicrurus forficatus) rear view, Berenty Reserve, Madagascar
Harvest mice (Micromys minutus), captive, UK, June
tail detail of Sperm Whale (Physeter macrocephalus) Pico Island, Azores, Portugal, Atlantic Ocean
Leopard (Panthera pardus) back of head, Samburu National Reserve, Kenya
Two Galapagos marine iguanas (Amblyrhynchus cristatus) on rock, Espanola Island, Galapagos, Vulnerable species
Portrait of Whale shark (Rhincodon typus) at the surface. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
Speckled bush-cricket (Leptophyes punctatissima) on fern fronds. Dorset, UK, July
Eurasian Lynx (Lynx lynx) adult sitting on boulder covered in deep snow in beech (Fagus sylvatica forest, captive in enclosure of the Bavarian Forest National Park, Germany, February
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
Black Swallowtail Butterfly (Papilio polyxenes), female on Cowpen Daisy / Golden Crownbeard (Verbesina encelioides). Laredo, Webb County, Texas, USA, April
Eurasian Sparrowhawk (Accipiter nisus) rear view perching on fence post with tail feathers spread. Vosges, France, January
Two African lion cubs (Panthera leo) playing on a road in the Kalahari Desert, Botswana
A pair of Ravens (Corvus corax) perched in bleached out pine tree, Estonia, March 2008
Confrontation between a great egret (Ardea alba) and grey heron (Ardea cinerea), Lake Csaj, Kiskunsagi National Park, Hungary
Hectors dolphin (Cephalorhynchus hectori) Akaroa Harbour, South Island, New Zealand, November