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Brown Roan Italian Spinone puppy, Riley, 13 weeks, chewing his tail
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Himalayan monal (Lophophorus impejanus) male perched on branch, Uttarakhand, India
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Green Sea Turtle (Chelonia mydas) returning to sea after Egg laying, Raine Island, Australia. Raine Island is the largest and most important green sea turtle nesting area in the world, with over 14
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Grey heron (Ardea cinerea) beneath waterfall. Ambleside, Lake District, UK, November. Highly commended, Habitat category, British Wildlife Photography Awards (BWPA) competition 2012
African forest elephant (Loxodonta cyclotis) in water, Lekoli River, Republic of Congo (Congo-Brazzaville), Africa. Vulnerable species
Gemsbok (Oryx gazella) female with two calves, Namibrand Reserve, Namib Desert, Namibia, Nambia
Sable Border Collie puppy, 9 weeks, looking over his shoulder
Yellowhammers (Emberiza citrinella) coming in to land of a sheaf of oats in winter. Perthshire, Scotland, February
Southern stingray (Hypanus americanus) swimming over sand ripples on sandbar, Stingray City, Grand Cayman. British West Indies
RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
Red fox (Vulpes vulpes) rear view sitting on grass close to its den, Derbyshire, UK
Baltimore oriole (Icterus galbula) female perched on Berlandiers fiddlewood (Citharexylum berlandieri). South Padre Island, Texas, USA. April
Ocelot (Leopardus pardalis) in rainforest, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Red deer stag (Cervus elaphus) rear view, shaking water off itself after the rain, Bradgate Park, Leicestershire, UK. GDT European Wildlife Photographer of the Year Competition 2021 Winner Mammal
Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Golden-fronted woodpecker (Melanerpes aurifrons), male, Cozad Ranch, Rio Grande Valley, Texas, USA, March
Western tanager (Piranga ludoviciana), male perched on aspen branch in spring, Mono Lake Basin, California, USA, May
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Golden pheasant (Chrysolophus pictus) male walking through the forest, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China. Endemic species for China
Welsh mountain pony (Equus caballus) with a view of the beach and Bell heather (Erica cinerea), St. Davids Head, Pembrokeshire Coast National Park, Pembrokeshire, Wales, UK, July
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Rear view of Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May
Brown Pelican (Pelecanus occidentalis) flying above the waves. Isabela Island, Galapagos Islands, Ecuador
Hoatzin (Opisthocomus hoazin) Yasuni National Park, Orellana, Ecuador
Female Kestrel (Falco tinnunculus) perched in the entrance to a nest hole in an old tree, Hertfordshire, England, UK, June
Tibetan antelope (Pantholops hodgsonii) herd in the early morning with a low moon, Kekexili, Qinghai, Tibetan plateau, China, January
Roan antelope (Hippotragus equinus) with young offspring, Mlilwane nature reserve breeding programme, Swaziland, Southern Africa
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Three Ring-tailed Lemurs (Lemur catta) sun basking at dawn. Berenty Private Reserve, southern Madagascar. (digitally stitched image)
RF- Rear view of male Bengal tiger (Panthera tigris tigris) walking on track. Bandhavgarh National Park, India. Endangered species
Barn owl (Tyto alba) perched at entrance to tree hollow, Hampshire, UK. October. Captive, controlled conditions
Yellow fiddler crab (Uca sp. ) portrait, Sunderban tiger reserve, West Bengal, IndiaYellow fiddler crab (Uca sp.) portrait, Sunderban tiger reserve, West Bengal, India
Sequence of tail fluke of Sperm whale (Physeter macrocephalus) diving. Dominica. Caribbean Sea, Atlantic Ocean. Digital composite
Common kingfisher (Alcedo atthis) female, perched on frosty branch in winter, Bavaria, Germany. January
Puma (Puma concolor) female, stalking Guanaco (Lama guanicoe) herd in snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Spotted flycatcher (Muscicapa striata) perched on branch in morning sunlight, Grazalema, Andalusia, Spain. August
Eurasian kingfisher (Alcedo atthis) perched on branch in autumn, Sierra de Grazalema Natural Park, Andalusia, southern Spain. November
Humpback whale (Megaptera novaeangliae) rear view of males escort, with adult female and male calf in the background. Vava'u, Tonga, South Pacific
Stork-billed kingfisher (Pelargopsis capensis) perched on branch with beak open, Bardia National Park, Terai, Nepal
Red grouse (Lagopus lagopus) on moorland at sunrise, Peak District National Park, UK. August