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Red squirrel (Sciurus vulgaris) amongst Bell heather (Erica cinerea) Cairngorms National Park, Highlands, Scotland, August
Red Squirrel (Sciurus vulgaris) male standing alert, Cairngorms National Park, Highlands, Scotland, UK, May
Red deer stag (Cervus elaphus) in rugged habitat standing next to old gnarled Scots pine tree, Western Highlands, Scotland, UK, December
Red squirrel (Sciurus vulgaris) close-up climbing down tree, Cairngorms National Park, Highlands, Scotland, UK, April
Pine marten (Martes martes) on branch of Scots pine tree. Perthshire, Highlands, Scotland, UK, May
Red Squirrel (Sciurus vulgaris) close-up portrait of red squirrel sniffing. Cairngorms National Park, Highlands, Scotland, UK, March
Scots pine (Pinus sylvestis) cone that been partially chewed / eaten by a Red squirrel (Sciurus vulgaris) on lichen covered forest floor. Cairngorms National Park, Highlands, Scotland, UK, August
Chickweed wintergreen ( Trientalis europaea) flower, Cairngorms National Park, Scotland, UK, June
Scots Pine forest (Pinus sylvestris) with early morning misty light filtering through the forest. Cairngorm National Park, Highlands, Scotland, UK, October 2014
Scots pine (Pinus silvestris) and Silver birch (Betula pendula) forest, with natural regeneration of pine and birch seedlings. Glen Strathfarrar, Highlands, Scotland, UK
Scots pine (Pinus sylvestris) seedling growing amongst lichens and heather. Cairngorms National Park. Highlands, Scotland, September
Red Squirrel (Sciurus vulgaris) foot prints in snow, Cairngorms National Park, Highlands, Scotland, UK, January
Scots pine (Pinus sylvestris) forest looking skywards towards forest canopy. Cairngorms National Park, Highlands, Scotland, UK, December
Twinflower (Linnaea borealis) in pine woodland, Cairngorms National Park, Highlands, Scotland, UK, June
Wet tree bark of Scots pine tree (Pinus sylvestris) Cairngorms National Park, Highlands, Scotland, UK, November
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
White Water Lily (Nymphaea alba) in flower, Cairngorms National Park, Scotland, UK
Scots pine (Pinus sylvestris) forest in evening light, Abernethy, Cairngorms National Park, Scotland, UK, June
Red squirrel (Sciurus vulgaris) on tree trunk, Cairngorms National Park, Scotland, UK, August
Night sky over Loch Morlich with star trails, Cairngorms National Park, Cairngorms, Scotland, UK, September 2015
Autumn colours and stormy clouds reflected in the calm waters of Loch Insh, Cairngorms National Park, Scotland, UK, November 2015
Iceberg in the sea outside the Kangia Ilulissat Icefjord. Iceberg from the Sermeg Kujalleg Glacier, UNESCO World Heritage Site, Kalaallit Nunaat, Greenland, August 2014
Nyiragongo volcano lava lake, Virungas National Park, Democratic Republic of Congo February 2015
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Black rhinoceros (Diceros bicornis) female and young in the mud, Masai-Mara Game Reserve, Kenya. February
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
The Pont St-Benezet, Palais des Papes and Rhone River at dusk, Avignon, Provence, France, October 2012
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Ground squirrels (Xerus inauris ) one with its tail over the back of another, Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Pair of Blesbok (Damaliscus pygargus) galloping across the open flood plains of the Bushmans River, Kariega Game Reserve, South Africa. Overall winner of the GDT Competition 2014
African fish eagle (Haliaeetus vocifer) in flight, Chobe National Park, Botswana
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in March. Wapusk National Park. Churchill, Manitoba, Canada
Masai Giraffe (Giraffa camelopardalis tippelskirchi) juveniles, Masai Mara Game Reserve, Kenya
Wood Anemones (Anemone nemorosa) in flower on woodland floor, low angle fish eye view, Peak District National Park, Derbyshire, UK, May
Pika (Ochotona princeps) in scree rock pile, Sheepeaters Cliff, Yellowstone National Park, Wyoming, USA, September
American Red fox (Vulpes vulpes fulva) baby leaping on its disinterested mother. Grand Teton National Park, Wyoming, USA, May
Himalayan Blue sheep (Pseudois nayaur) Hemas National Park, Ladakh, India
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Lion (Panthera leo) male smelling, flehmen response, Masai-Mara Game Reserve, Kenya. Vulnerable species
White rhino (Ceratotherium simum) female portrait, portrait, Nakuru National Park, Kenya
Lily of the valley (Convallaria majalis), Foret de Puvenelle, Lorraine Regional Natural Park, Lorraine, France, May
Female Pintail (Anas acuta) with chicks, Katmai National Park and Preserve, Alaska, USA, July
Common juniper (Junipera communis) and Common beech (Fagus sylvatica) trees growing on pasture land grazed by livestock in summer on subalpine meadow
Red-vented bulbul (Pycnonotus cafer) Bandhavgarh National Park, India, March
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May