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Marbled white butterfly (Melanargia galathea) roosting among tall grasses at sunrise and bathed in warm, early morning sunlight, Volehouse Moor, Devon, UK. July
RF - Goshawk (Accipiter gentilis) juvenile on snow covered branch. Flatanger, Trondelag, Norway. February. (This image may be licensed either as rights managed or royalty free.)
Black woodpecker (Dryocopus martius) male on tree trunk, portrait. Kuusamo, Northern Ostrobothnia, Finland. February
Common red soldier beetle (Rhagonycha fulva) climbing up grass with dew drops, Hertfordshire, England, UK, July - Focus Stacked
Zebra / leopard shark (Stegostoma fasciatum) swimming over the reef, Vaavu atoll, Maldives, Indian Ocean
Grey heron (Ardea cinerea) immature portrait. London, England, UK, December
Fox (Vulpes vulpes) in snow, Londong, England, UK, January
Robin (Erithacus rubecula) in snow, Broxwater, Cornwall, UK. March
Egyptian vulture (Neophron percnopterus) adult with the Haggeher mountains in the background, Socotra Island, Yemen, March
Great blue heron (Ardea herodias) with hatchling sea turtle prey, Las Bachas, Santa Cruz Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) on head of Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) drinking, Alcedo Volcano, Isabela Island, Galapagos
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Anolis lizard (Anolis sp. ) male displaying in Corcovado National Park, Costa RicaAnolis lizard (Anolis sp.) male displaying in Corcovado National Park, Costa Rica
Copper-cheeked frog (Chalcorana labialis) from riparian lowland rainforest on the Cross-island trail, Tioman Island, Malaysia
Potato grouper (Epinephelus tukala) in Passe Dubois / Dubois channel, Aldabra, Indian Ocean
RF-Polar bear (Ursus maritimus) swimming in Beaufort Sea, portrait. Near Kaktovik, Arctic National Wildlife Refuge, Alaska, USA. September
Corncrake (Crex crex) calling at dawn on machair, North Uist, Outer Hebrides, Scotland, UK, June
Dipper (Cinclus cinclus) in river, Finland, March
Red-throated Diver (Gavia stellata) portrait, Iceland, June
Hazel Grouse (Tetrastes bonasia), Kuusamo, Finland, December
Mediterranean gull (Larus melanocephalus) in summer plumage, Hayling Island, Hampshire, England, UK, July
Grey shrike (Lanius meridionalis) perched, Navarra province. Spain, July
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
RF- Verreauxs sifaka lemur (Propithecus verreauxi) hanging from branch while feeding in canopy. Berenty Private Reserve, southern Madagascar. Endangered species
Atlantic walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks, Svalbard, Norway, June
Short-snouted seahorse (Hippocampus hippocampus) portrait. Tenerife, Canary Islands
Mandarin duck (Aix galericulata) reflected in water. Richmond Park, London, England, UK
Cheetah (Acinonyx jubatus) walking, silhouetted at dusk. Zimanga private game reserve, KwaZulu-Natal, South Africa
Ruwenzori side-striped chameleon, (Trioceros rudis), hanging from fern, Nyungwe Forest NP, Rwanda, December. Non-ex
Snowshoe hare (Lepus americanus) adult with coat changing into summer colors, south side of the Brooks Range, Alaska, USA
Sardinian warbler (Sylvia melanocephala) Sierra de Andujar Jaen Andalucia, Spain, April
California Quail (Callipepla californica), adult male perched on bush, acting as sentinel, Montana De Oro State Park, California, USA, July
Coffee bee hawk moth (Cephaonodes hylas) Amagasaki City, Hyogo, Japan
African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Many-coloured rush tyrant (Tachuris rubrigastra) perched on reed stem at the waters edge. Lake Titicaca, Puno, Peru. September. Cropped
Onager (Equus hemionus), Negev desert, Israel, April
Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June
Blue-headed quail-dove (Starnoenas cyanocephala) profile, Cuba. Endangered species. Endemic
Male Reindeer (Rangifer tarandus) on autumn tundra. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, October
Black bear (Ursus americana) foraging for alpine berries during Autumn, Cascade Mountains, Washington, USA
Greenfinch (Carduelis chloris) on blackthorn / sloe berries, UK
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Violet tailed sylph (Aglaiocercus coelestis) Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Sword billed hummingbird (Ensifera ensifera) profile showing beak is longer than body, Yanacocha Reserve, Jocotoco Foundation, 3, 200m altitude on west slope of Pichincha Volcano
Masked trogon (Trogon personatus) with insect prey, Bellavista cloud forest private reserve, 1700m altitude, Tandayapa Valley, Andean cloud forest, Ecuador
Purple gallinule (Porphyrio martinicus) reaches for giant bulrush (Schoenoplectis californicus) seeds. Wakodahatchee Wetlands, Florida, USA, April