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White throated kingfisher (Halcyon smyrnensis) perched on a reed, Sunderban tiger reserve, West Bengal, India
Caribbean flamingo (Phoenicopterus ruber) two walking, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September. Bookplate
Elephant (Loxodonta africana) female and calf under tree in the rain, Masai-Mara game reserve, Kenya
Oystercatcher (Haematopus ostralegus) in flight over wildflowers, Great Saltee, Saltee Islands, County Wexford, Republic of Ireland. May
Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia
Roe deer (Capreolus capreolus) on a misty morning at the edge of a pond. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. December
Mute swan (Cygnus olor) on river in evening light, Gelderse Poort, near Nijmegen, The Netherlands. June
Banded demoiselle (Calopteryx splendens), male and female silhouetted on reed, Lower Tamar Lakes, Cornwall, UK. June
Kingfisher (Alcedo atthis) diving into water to catch fish, Near Bourne, Lincolnshire, England, UK. January
RF - Female Banded demoiselle damselfly (Calopteryx splendens) roosting among grasses. Cornwall, UK. July. (This image may be licensed either as rights managed or royalty free)
RF -Two King penguins (Aptenodytes patagonicus) head to head, South Georgia, South Atlantic. (digitally stitched image) (This image may be licensed either as rights managed or royalty free.)
Grey wolf (Canis lupus) standing among trees in snow-covered forest, Finland. November
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Robin (Erithacus rubecula) perched on fallen apple in snow, Bishopswood, Somerset, UK. January
Moorhen (Gallinula chloropus) on water in falling snow, near Bourne, Lincolnshire, England, UK. March
Eurasian blue tit (Cyanistes caeruleus) perched on snow-covered grass, puffed up to keep warm, Białowieza National Park, Poland. January
Two European goldfinches (Carduelis carduelis) fighting over food in snow, Wrocław, Lower Silesia, Poland. February
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Mourning dove (Zenaida macroura) perched on branch in snow storm; Milford, Connecticut, USA. February
Collie standing in snowy, frozen marsh, Eagle River, Alaska, USA. April
RF - Atlantic puffin (Fratercula arctica) among sea campion (Silene maritima), Skomer Island, Pembrokeshire, UK. May
Two Grey wolves (Canis lupus) play fighting in snow, Finland. November
Ruby-tailed / Cuckoo wasp (Chrysis comparata) on Mexican daisy / Fleabane (Erigeron karvinskianus). Podere Montecucco, Orvieto, Italy. June
Saunders bee hawkmoth (Hemaris saundersii) resting on a leaf, Shogran, Khyber-Pakhtunkhwa, Pakistan
Cheetah (Acinonyx jubatus) standing on top of a rock, Spain. Captive
Cheetah (Acinonyx jubatus) silhouetted at sunset, Spain. Captive
Two Caracal (Caracal caracal) cubs, aged 9 weeks, walking over rocks, Spain. Captive, occurs in Africa and Asia
Caracal (Caracal caracal) male, standing on rock against blue sky, Spain. Captive, occurs in Africa and Asia
RF - Two Cheetahs (Acinonyx jubatus) standing and sitting on rocks against blue sky, Spain. Captive. (This image may be licensed either as rights managed or royalty free.)
RF - Two Caracal (Caracal caracal) cubs, aged 9 weeks, sitting on hind legs looking up, Spain. Captive, occurs in Africa and Asia
Leopard (Panthera pardus) female, walking across grassland in morning sunlight, Kenya
Grey-headed flying-fox (Pteropus poliocephalus) female, in flight carrying her young, Myuna Wetlands, Doveton, Victoria, Australia
Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp)Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp.), Brooks Falls, Katmai National Park, Alaska. July
Brown bear (Ursus arctos) female with two cubs walking through misty, woodland meadow, Finland. June
Ptarmigan (Lagopus mutus) in winter plumage, resting in snow, Svalbard, Norway. April
Polar bear (Ursus maritimus) female standing on snow with cub peering out behind, Svalbard, Norway. April
Grizzly bear (Ursus arctos) watching for Salmon at top of waterfall, Brooks Falls, Katmai National Park, Alaska, USA. July
Poppy bee (Osmia papaveris) resting on Common poppy (Papaver rhoeas) cutting piece of petal to use as nesting material, Bamberg, Germany. June
Blue and yellow macaw (Ara ararauna) in flight, Pantanal, Mato Grosso do Sul, Brazil
Giant anteater (Myrmecophaga tridactyla) walking, Pantanal, Mato Grosso do Sul, Brazil
Red and green macaw (Ara chloropterus) in flight, Burraco das Araras, Pantanal wetlands, Mato Grosso do Sul, Brazil
Red and green macaw (Ara chloropterus) head portrait, Burraco das Araras, Pantanal wetlands, Mato Grosso do Sul, Brazil
Great kiskadee (Pitangus sulphuratus) perched on a twisted branch, Pantanal wetlands, Mato Grosso, Brazil
Capybara (Hydrochoerus hydrochaeris) sitting in wetland vegetation with Giant cowbird (Molothrus oryzivorus) perched on its back, Cuiaba River, Pantanal, Mato Grosso, Brazil
Jaguar (Panthera onca) resting on branch, Cuiaba River, Pantanal wetlands, Mato Grosso, Brazil
Yacare caiman (Caiman yacare) with mouth open, head portrait, Cuiaba River, Pantanal, Mato Grosso, Brazil
Jaguar (Panthera onca) standing on riverbank, Cuiaba River, Pantanal wetlands, Mato Grosso, Brazil
