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Guianan brown capuchin (Sapajus apella) chewing on branch, Pampas del Yacuma Protected Area, Bolivia
Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January
Chimpanzee male (Pan troglodytes schweinfurthii) sleeping on the forest floor with feet up, Kibale National Park, Uganda. January
Tapanuli Orangutan (Pongo tapanuliensis). Togus, adult flanged male. Batang Toru Forest, Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
Maroon / Red leaf monkey / Langur (Presbytis rubicunda) eating fruits in tree, Danum Valley Conservation Area, Sabah, Borneo, Malaysia
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia
Ring tailed lemur (Lemur catta) backlit in late afternoon light, Berenty Private Reserve, southern Madagascar
Male Indri (Indri indri) leaping through the rain forest canopy. Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Coquerels sifaka (Propithecus coquereli) female with young on back climbing in bamboo, Anjajavy Private Reserve, north west Madagascar
Colombian spider monkey (Ateles fusciceps rufiventris) group. Captive, Critically endangered species
Tufted grey langur (Semnopithecus priam) baby in tree, Nilgiri Biosphere Reserve, Karnataka, India
Verreauxs Sifaka (Propithecus verreauxi) holding baby, Berenty Reserve, Madagascar
Eastern Lesser Bamboo Lemur (Hapalemur griseus) Andasibe-Mantadia National Park, Alaotra-Mangoro Region, eastern Madagascar
Hanuman Langurs (Semnopithecus entellus) in Bougainvillea Mandore Garden, Jodhpur, India
Golden-crowned Sifakas or Tattersalls Sifakas (Propithecus tattersalli) feeding in forests near Andranotsimaty, Daraina, north east Madagascar. Critically Endangered
Adult Red Ruffed Lemur (Varecia rubra) resting in rainforest canopy. Masoala National Park, north east Madagascar. Endangered
Male Indri (Indri indri) reaching for leaves / foraging in forest understorey. Mitsinjo Reserve, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Red-shanked Douc langur (Pygathrix nemaeus). Captive, occurs in Cambodia, Lao Peoples Democratic Republic and Vietnam. Endangered species
Mountain forests where the Yunnan snub-nosed monkey (Rhinopithecus bieti) lives, Yunnan, China
Crowned sifaka (Propithecus coronatus), Katsepy, Madagascar
Coquerels sifaka (Propithecus coquereli) curious individual up in tree, Ankarafantsika National Park, Madagascar
Golden-brown mouse lemur (Microcebus ravelobensis), Ankarafantsika National Park, Madagascar
Female Yunnan Snub-nosed monkey (Rhinopithecus bieti) feeding, Tacheng Yunnan snub-nosed monkey National Park, Yunnan, China
Golden lion tamarin {Leontopithecus rosalia} captive, Peru, Critically endangered
Tibetan macaque (Macaca thibetana) infant in tree, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Bornean orangutan (Pongo pygmaeus) mother and baby, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Orangutan (Pongo pygmaeus) adult male, Nyaru Menteng Orangutan Reintroduction Project, Central Kalimantan, Borneo, Indonesia
Orang utan (Pongo pygmaeus) head portrait of female, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Sarawak, Borneo, Malaysia, Endangered
Orang utan (Pongo pygmaeus) head portrait of dominant male Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Orang utan (Pongo pygmaeus) portrait, swinging, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Juvenile female Bornean Orangutan (Pongo pygmaeus) called Betsy (daughter of Beth) hanging from vine, Gunung Palung NP, Borneo
Japanese Macaque (Macaca fuscata) in snow, Jigokudani, Japan. February
Japanese Macaque (Macaca fuscata) Jigokudani, Japan, January
Japanese Macaques (Macaca fuscata) lined up in hotsprings, Jigokudani, Japan, February
Japanese Macaque (Macaca fuscata) in thermal pool, falling asleep, Jigokudani, Japan, January
Japanese macaque portrait {Macaca fuscata} Jigokudani, Japan
Tibetan macaque (Macaca thibetana) juveniles playing, hanging upside down in tree, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Phayres leaf monkey (Trachypithecus phayrei) siiting on a tree at He Xin Chang Forest reserve, Dehong Prefecture, Yunnan Province, China, May
Yunnan snub-nosed monkey (Rhinopithecus bieti) sitting in a tree at Ta Cheng Nature Reserve, Yunnan, China. October
Mantled howler monkey (Alouatta palliata) with bokeh, Costa Rica
White-faced capuchin monkey (Cebus capucinus) finding insects in wood, Curu National Park, Costa Rica. March 2015
Hanuman langur (Semnopithecus / Presbytis entellus), female with baby at water tapHanuman langur (Semnopithecus/ Presbytis entellus), female with baby at water tap, Ranthambhore National Park, Rajasthan, India
Black and white ruffed lemur (Varecia variegata variegata) hanging from branch, Vakona island, Andasibe area, Madagascar. Captive
Diademed sifaka (Propithecus diadema diadema) pair, Maromizaha Reserve, Andasibe Mantadia area, eastern Madagascar. August
RF- Black and white ruffed lemur (Varecia variegata variegata) sunbathing in the early morning, Vakona island, Andasibe area, Madagascar. Captive
Ankarana sportive lemur (Lepilemur ankaranensis), Ankarana National Park, Madagascar
Southern plains grey langur (Semnopithecus entellus / Presbytis entellus) tiny juvenile, Jodhpur, Rajasthan India
Southern plains grey / Hanuman langur {Semnopithecus dussumieri} drinking, Sariska NP, Rajasthan, India
