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Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Portrait of Waglers / Temple Pitviper (Tropidolaemus wagleri) showing the thermo-receptive pit organ between the eye and mouth. Controlled conditions. Mindanao, Philippines, February
Hawksbill turtle (Eretmochelys imbricata) ventral view, Philippines, Pacific Ocean. Critically endangered
Coral reef with Threadfin anthias (Pseudanthias huchtii) shoal with Bluegreen chromis (Chromis viridis) amongst them, under shallow sea with a coconut tree forest in background, Philippines
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
Honeycomb / Cushion starfish (Pentaceraster alveolatus) composite image on black background showing colour variations Malapascua Island, Philippines, Indo-Pacific species
Venomous Yellow-lipped sea krait (Laticauda colubrina) heading up to the surface to breathe, Philippines
Blackspotted puffer (Arothron nigropunctatus) curled up on the reef for the night, with a transparent cleaner shrimp (Urocaridella antonbrunii) inspecting its tail, Philippines
A massive school of millions of sardines (Sardina pilchardus), Moalboal, Cebu, Central Visayas, Philippines
Ornate ghost pipefish (Solenostomus paradoxus), juvenile, hides within a crinoid, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Pair of Coleman shrimps (Periclimenes colemani) living in Fire urchin (Asthenosoma varium). The female is the larger shrimp. Anilao, Batangas marine protected area, Luzon, Philippines
Wide angle macro photo of an algae octopus (Abdopus aculeatus) as it reaches out and investigates the camera. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Elegant conch (Lentigo pipus) close up of eye stalks, Philippines
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Coral reef scene, with Philippines chromis (Chromis scotochiloptera), hard coral (Acropora sp.) and purple elephant ear sponges (Ianthella basta)
Leopard shark (Stegostoma fasciatum) cruising along the edge of an underwater drop off, Philippines
Emperor shrimp (Zenopontonia rex) female rides on the back of a nudirbranch (Tambja morosa), while picking food from the seabed. Anilao, Batangas marine protected area, Luzon, Philippines
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Ring-tailed / Golden cardinalfish (Ostorhinchus aureus) male incubating its eggs in its mouth, Sulu sea, Philippines
Stargazer (Uranoscopus sulphureus) half buried in the sand, Sulu sea, Philippines
Henshaws snake eel (Brachysomophis henshawi) burried in the sand, Sulu sea, Philippines
Portrait of a Thorny seahorse (Hippocampus histrix) near Malapascua, Philippines
Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines
Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp). Anilao, Philippines
Close up detail of the rhinophores and caruncle of a Nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
False clown anemonefish (Amphiprion oceallaris) guarding their eggs laid on a coral rock beneath a sea anemonePhilippines. Bohol Sea, Tropical West Pacific Ocean
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines
Bargibants pygmy seahorse (Hippocampus bargibanti) wrapped around Eye fan coral (Cnidaria sp). Philippines
Bargibant's pygmy seahorse (Hippocampus bargibanti) wrapped around Eye fan coral (Cnidaria sp). Philippines
Honeycomb / Cushion starfish (Pentaceraster alveolatus) Malapascua Island, Philippines, September
Emperor shrimp (Periclemenes imperator) on sea cucumber (Thelenota ananas) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Longnose hawkfish (Oxycirrhites typus) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Amphipod (Amphipoda) living inside sea squirt (Rhopalaea sp. ) Anilao, Batangas, Luzon, PhilippinesAmphipod (Amphipoda) living inside sea squirt (Rhopalaea sp.) Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Long-beaked Coralfish (Chelmon rostratus) eye, Inanuran Island, Danajon Bank, Central Visayas, Philippines, April
Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply
Harlequin swimming crab (Lissocarcinus laevis) sheltering beneath the tentacles of a tube anemone (Cerianthus sp.). Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
Soft coral crab (Hoplophrys oatesil) climbing out onto a branch of yellow and red soft coral (Dendronephthya sp.) at night. Anilao, Batangas, Luzon, Philippines