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RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Colorful Pfeffers flamboyant cuttlefish (Metasepia pfefferi) swimming over a colony of cup coral at night, Philippines
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Larval Wonderpus octopus (Wunderpus photogenicus) drifting in the open ocean at night off Anilao, Philippines
False tiger moth (Dysphania militaris) Isle of Marinduque, Philipines
African pompano (Alectis ciliaris) juvenile with long trailing dorsal and anal fins, swimming in mid-water at night. In-camera double exposure
Jade vine (Strongylodon macrobotrys) in glasshouse at Kew Gardens, London, UK. Endemic to the Philippines. Pollinated by sunbirds
Moorish idol (Zanclus cornutus) juvenile with transparent skin in the open ocean at night off Anilao, Philippines
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Wonderpus octopus (Wunderpus photogenicus) in its juvenile or larval stage. At this stage it is transparent and drifts in the open ocean as plankton. Anilao, Philippines
Trumpetfish (Aulostomus chinensis) on the coral drop off, Sulu Sea, Philippines
Clownfish (Amphiprion sp) in anemone home, Philippines
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Variegated feather duster (Bispira variegata) Sabellidae. Puerto Galera, Philippines
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
Pfeffers flamboyant cuttlefish (Metasepia pfefferi) composite image on black background showing colour variations, Puerto Galera, Philippines, Indo-Pacific
Scalloped ribbonfish (Zu cristatus) juvenile, Balayan Bay, Batangas, the Philippines
The Chocolate Hills are a geological formation in Bohol province in the Philippines. More than a thousand of these hills occur in a 50 square kilometre area
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Paper nautilus (Argonauta hians) in open water at night. Anilao, Batangas marine protected area, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Kaleidoscopic image of Variegated feather duster (Bispira variegata), Sabellidae. Puerto Galera, Philippines
Halimeda ghost pipefish (Solenostomus halimeda) mimicking the halimeda algae. Sulu sea, Philippines
False clown anemonefish (Amphiprion ocellaris) in a magnificent sea anemone (Heteractis magnifica), Sulu sea, Philippines
A juvenile soldierfish, possibly a Crescent-tail bigeye (Priacanthus hamrur) in the open ocean at night off Anilao, Philippines
Larval flounder (possibly Laeops Kitahara) in the open ocean at night off Anilao, Philippines
Pygmy seahorse (Hippocampus bargibanti) holding on to a branch of a sea fan coral. Malapascua, Philippines
Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp). Anilao, Philippines
Rainforest and river outside the Batak village of Sitio Manggapin in Cleopatra sRainforest and river outside the Batak village of Sitio Manggapin in Cleopatras Needle Critical Habitat, Palawan, Philippines
Close up detail of the rhinophores and caruncle of a Nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
RF - Orange skunk clownfish (Amphiprion sandaracinos) in anemone, Puerto Galera, Philippines
Magnificent feather duster worm (Protula magnifica) Puerto Galera, Philippines
Goldring tunicate (Clavelina robusta) Puerto Galera, Philippines
Hawksbill turtle (Eretmochelys imbricata) ventral view, Philippines, Pacific Ocean. Critically endangered
Coral reef with Threadfin anthias (Pseudanthias huchtii) shoal with Bluegreen chromis (Chromis viridis) amongst them, under shallow sea with a coconut tree forest in background, Philippines
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
Honeycomb / Cushion starfish (Pentaceraster alveolatus) composite image on black background showing colour variations Malapascua Island, Philippines, Indo-Pacific species
Venomous Yellow-lipped sea krait (Laticauda colubrina) heading up to the surface to breathe, Philippines
Blackspotted puffer (Arothron nigropunctatus) curled up on the reef for the night, with a transparent cleaner shrimp (Urocaridella antonbrunii) inspecting its tail, Philippines
A massive school of millions of sardines (Sardina pilchardus), Moalboal, Cebu, Central Visayas, Philippines
Ornate ghost pipefish (Solenostomus paradoxus), juvenile, hides within a crinoid, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Pair of Coleman shrimps (Periclimenes colemani) living in Fire urchin (Asthenosoma varium). The female is the larger shrimp. Anilao, Batangas marine protected area, Luzon, Philippines
Wide angle macro photo of an algae octopus (Abdopus aculeatus) as it reaches out and investigates the camera. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Elegant conch (Lentigo pipus) close up of eye stalks, Philippines
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Coral reef scene, with Philippines chromis (Chromis scotochiloptera), hard coral (Acropora sp.) and purple elephant ear sponges (Ianthella basta)
Leopard shark (Stegostoma fasciatum) cruising along the edge of an underwater drop off, Philippines
Emperor shrimp (Zenopontonia rex) female rides on the back of a nudirbranch (Tambja morosa), while picking food from the seabed. Anilao, Batangas marine protected area, Luzon, Philippines
Ring-tailed / Golden cardinalfish (Ostorhinchus aureus) male incubating its eggs in its mouth, Sulu sea, Philippines
Stargazer (Uranoscopus sulphureus) half buried in the sand, Sulu sea, Philippines
Henshaws snake eel (Brachysomophis henshawi) burried in the sand, Sulu sea, Philippines
Portrait of a Thorny seahorse (Hippocampus histrix) near Malapascua, Philippines
Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines
False clown anemonefish (Amphiprion oceallaris) guarding their eggs laid on a coral rock beneath a sea anemonePhilippines. Bohol Sea, Tropical West Pacific Ocean
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines
Bargibants pygmy seahorse (Hippocampus bargibanti) wrapped around Eye fan coral (Cnidaria sp). Philippines
Honeycomb / Cushion starfish (Pentaceraster alveolatus) Malapascua Island, Philippines, September
Broadclub cuttlefish (Sepia latimanus) in translucent egg casing on the reef, Philippines, Asia
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Philippine eagle owl (Bubo philippinensis) captive, endemic to the Philippines
Thimble jellyfish (Linuche unguiculata) aggregation, My Paradise Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Ascidian (Rhopalaea sp) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Longnose hawkfish (Oxycirrhites typus) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Spaghetti garden eels (Gorgasia maculata) stretching up out of their burrows on a rubble slope, with Mysiid shrimp swimming around their bodies near the seabed. Dumaguete, Dauin, Negros, Philippines
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Spotted jellyfish (Mastigias papua) Inanuran Island, Danajon Bank, Central Visayas, Philippines, April
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
Aggregation of tiny isopods (Santia sp)Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Portrait of Waglers / Temple Pitviper (Tropidolaemus wagleri) showing the thermo-receptive pit organ between the eye and mouth. Controlled conditions. Mindanao, Philippines, February
Emperor shrimp (Periclemenes imperator) on sea cucumber (Thelenota ananas) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
