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Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Portrait of Male Sockeye salmon (Oncorhynchus nerka) showing teeth. Adams River, British Columbia, Canada, October
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
Whitetip shark (Carcharhinus longimanus), accompanied by Pilotfish (Naucrates ductor), swims towards the camera as it cruises beneath the surface of the Red Sea, close to Little Brother Island
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Yellowhead jawfish (Opistognathus aurifrons) spitting out sand as it digs its burrow. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
DELETE Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea
RF - Spinyhead blenny fish (Acanthemblemaria spinosa) extends out of its hole in a red sponge to snare passing food. Caribbean Sea
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Florida manatee (Trichechus manatus latirostrus) with Blue gill sunfish (Lepomis macrochirus) cleaning it, in a freshwater spring, beneath trees. Three Sisters Spring, Crystal River, Florida, USA
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Garibaldi fish (Hypsypops rubicundus) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphere Reserve, Baja California, Mexico, May
Sardine bait ball (Sardinops sagax) escaping Striped marlin (Tetrapturus audax) that is attempting to feed on them. West Coast of Baja California Peninsula, Mexico. Pacific Ocean
Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp)Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp.), Brooks Falls, Katmai National Park, Alaska. July
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp. ) hunting a large school of Sardines (Sardina sp)Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp.) hunting a large school of Sardines (Sardina sp.), Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Two Blue spotted mud skippers (Boleophthalmus boddarti) fighting at water's edge, Sunderban tiger reserve, West Bengal, India
Coral reef with Threadfin anthias (Pseudanthias huchtii) shoal with Bluegreen chromis (Chromis viridis) amongst them, under shallow sea with a coconut tree forest in background, Philippines
Shoal of Lowfin chub (Kyphosus vaigiensis) evading a Coral grouper (Cephalopholis miniata), Tubuai, French Polynesia, Pacific Ocean
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Bigeye trevallies (Caranx sexfasciatus) shoal. Darwin Island, Galapagos National Park, Pacific Ocean
Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp)Common ghostgoby (Pleurosicya mossambica) camouflaged on a hard coral colony (Plesiastrea sp.), Ambon Bay, Maluku Archipelago, Indonesia. Banda Sea
Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean
Healthy coral reef landscape, Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
Scrapemouth mbuna cichlid (Labeotropheus trewavasae) swimming. Thumbi West island, Lake Malawi, Malawi
Small green utaka cichlid (Nyassachromis leuciscus) swimming. Thumbi West island, Lake Malawi, Malawi
