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Harestail cottongrass (Eriophorum vaginatum) in flower, Flow Country, Forsinard, Caithness, Highland, Scotland, UK, May
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Adder (Vipera berus) portrait coiled up, Powerstock Common nature reserve, Dorset, England, UK June
Secretary blenny (Acanthemblemaria maria) making a home in a deserved worm tube of a great star coral (Montastraea cavernosa), East End, Grand Cayman, Cayman Islands, British West Indies
Jackdaws (Corvus monedula) and Rooks (Corvus frugilegus) mixed winter flock taking flight from a frost covered field of crop stubble, Derbyshire, UK, January (non-ex)
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Great yellow gentian (Gentiana lutea) leaves. Haute-Savoie, Massif des Bauges, French Alps, May
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
Abstract reflection of reeds in rippled water, Westhay Moor SWT reserve, Somerset Levels, Somerset, England, UK, June 2011
Palm Frond detail. Mahahual Penninsula, South Yucatan Peninsula, Mexico
Eastern green mamba {Dendroaspis angusticeps} skin detail, captive, from East Africa
Detail of sponge (Chalinula nematifera) Maldives, Indian Ocean
Broken ice abstract, Jokulsarlon glacier lagoon. Southern Iceland, Iceland, Europe, March 2011
Detail in frozen waterfall. Selijalandfoss waterfall, Southern Iceland, Europe, March 2011
Masai Giraffe (Giraffa camelopardalis tippelskirchi) male and female (back). Masai-Mara Game Reserve, Kenya
Broad snouted caiman (Caiman latirostris) hatching from egg in nest, Sante Fe, Argentina, February
Various rockpool animals and seaweeds, photographed on a white background. Isle of Skye, Inner Hebrides, Scotland, UK, March. Digital composite
Snowflake magnified under microscope, Lilehammer, Norway
Aerial view saltmarshes of Bahia de Cadiz Natural Park, Huelva, Andalucia, Spain, March 2008
Aerial image showing sandbanks in the Bahamas archipelago, Caribbean, February 2012
Aerial view of Everglades National Park, Florida, USA, February 2012
Sand dunes in the Libyan desert, Sahara, Libya, North Africa, December 2007
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Bigeye Scad (Selarcrumenophthalmus). Misool, Raja Ampat, West Papua, Indonesia
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Looking up at Beech wood canopy (Fagus sylvatica) in autumn, Peak District National Park, Derbyshire, UK, November
Aerial view of Lake Natron, Rift Valley, Tanzania, Africa, August 2009
A lone Caribbean reef shark (Carcharhinus perezi) cruises over sand ripples. Walkers Cay, Northern Bahama Islands, Republic of Bahamas
Detail in ice pattern formations, the Netherlands
Sand dunes in the Libyan desert at dawn, Sahara, Libya, North Africa, November 2007
Autumn colours reflected in river water, Ontonagon River, flows into Lake Superior. Michigan, USA
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Aerial image showing sandbanks and islands in the Bahamas archipelago, Caribbean, February 2012
Scarlet ibis (Eudocimus ruber) roosting in trees on a small mangrove island in the Caroni Swamp, Caroni Bird Sanctuary, Trinidad
Fallen leaves and swirling foam patterns in River Affric, Glen Affric NNR, Cairngorms NP, Highlands, Scotland, UK, October
Abstract details of autumnal colour and trees. Subject motion created by moving the camera during exposure. Beacon Hill Country Park, The National Forest, Leicestershire, UK. November
Details of gills Fly Agaric fungus {Amanita muscaria} in autumn, Beacon Hill Country Park, The National Forest, Leicestershire, UK. November
Close-up of gills of mushroom, Bradgate Park, The National Forest, Leicestershire, UK, November
Ink cap fungus {Coprinus acuminatus} in taiga woodland, autumn, Laponia / Lappland, Finland
Parasol mushroom cap close-up detail {Macrolepiota procera} Europe
Common spangle galls of the Gall wasp (Neuroterus quercusbaccarum) on underside of oak leaf, Dorset, UK, September
Aerial view of saltmarshes and Brancaster and Scolt Head Island, Norfolk, UK, September
Female Adder (Vipera berus) basking in hazy September sun, Surrey, England
Orange Tip Butterfly (Anthocharis cardamines) female resting on bluebell, Lanhydrock woodland, Cornwall. May 2012