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Female Nursery web spider (Pisaura mirabilis) carrying egg sac, Peak District National Park, Derbyshire, UK. June
Female Leopard (Panthera pardus) carrying cub in her mouth to their new den, Kgalagadi Transfrontier Park, South Africa
Female Kingfisher (Alcedo atthis) covering chicks, aged one day, with her wings in an artificial nest, Italy
Male Kingfisher (Alcedo atthis) sitting on eggs for his brooding period in an artificial nest, Italy
Male Kingfisher (Alcedo atthis) feeding chicks, aged 9 days, with a large fish in artificial nest, Italy
Mother White-tailed deer (Odocoileus virginianus) allowing young to suckle, Finland. September
Adult Swift foxes (Vulpes velox) caring for pup at den, Montana, USA. June
Polar bear (Ursus maritimus) mother and cubs rubbing themselves against mounds of snow, Robeson Channel, North Greenland. August
Male Dendtric jawfish (Opistognathus dendriticus) with incubating eggs in his mouth, Dampier Strait, Raja Ampat, Indonesia, Pacific Ocean
Blue-footed booby (Sula nebouxii), incubating eggs with webbed feet, Seymour Island, Galapagos, South America
Swallow-tailed gull (Creagrus furcatus), pair guarding small chick, Genovesa Island, Galapagos, South America
Brown pelican (Pelecanus occidentalis) pair in breeding plumage, collecting nesting material, Santa Cruz Island, Galapagos, South America
Fishing cat (Prionailurus viverrinus) with two kittens, age 4 weeks, learning to hunt for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss
White stork (Ciconia ciconia) flying back to its nest in Oak (Quercus robur) tree to feed its three large chicks, Knepp Estate, Sussex, UK, June
Adult White stork (Ciconia ciconia) standing beside three large chicks on their nest, Knepp Estate, Sussex, UK, June
Panamic sergeant major (Abudefduf troschelii) guarding egg mass, removing seastar that was coming to close to egg mass, Punta Resbaladilla, south of El Tecolote, Gulf of California (Sea of Cortez)
Giant damselfish (Microspathodon dorsalis) guarding egg mass, Coronado Island, Loreto Bay National Park, Gulf of California (Sea of Cortez), Mexico, August
Coyote (Canis latrans) female with newborn pups. Yellowstone National Park, Wyoming, USA
Arboreal tarantula (Lampropelma sp. ) female guarding spiderlings at entrance to hollowArboreal tarantula (Lampropelma sp.) female guarding spiderlings at entrance to hollow retreat, 4 metres up a tree. Danum Valley, Sabah, Borneo
Swift fox (Vulpes velox) female cleans the fur of one of her kits while another tries to nurse, Eastern Colorado, USA, May
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Barbary macaque (Macaca sylvanus) baby sheltering between arms of mother and another adult. Gibraltar Nature Reserve, Gibraltar. August
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar
Australian sea lion (Neophoca cinerea) pup suckling. Kangaroo Island, South Australia. November
Mountain gorilla (Gorilla beringei) mother with one month baby feeding in tree. Member of Katwe group. Bwindi Impenetrable Forest National Park, Uganda, Africa
Red fox (Vulpes vulpes) female, suckling cubs age five weeks, the Netherlands
RF- Eurasian brown bear (Ursus arctos) mother walking with cub, Suomussalmi, Finland. July
Baby African elephant (Loxodonta africana), climbing up a riverbank, Chobe National Park, Botswana
Bengal Tiger (Panthera tigris tigris) mother and cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle
Australian sealion with young at sea surface {Neophoca cinerea} Australia. Endangered species