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Waved albatross (Phoebastria irrorata) chick, aged two weeks, begging next to parent on nest. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Pair of Swallow-tailed gulls (Creagrus furcatus) at nest, one incubating. Genovesa Island, Galapagos Islands, Ecuador
Blessed poison frog (Ranitomeya benedicta) male, carrying tadpoles on its back, transporting them to water, Yurimaguas, Alto Amazonas, Peru
Great crested grebe (Podiceps cristatus) carring three chicks on its back, Valkenhorst nature reserve, Valkenswaard, The Netherlands
Two banded plover (Charadrius falklandicus) female, incubating eggs on nest on pebbly beach, Cerro Avanzado protected area, Chubut, Patagonia, Argentina
Leatherback turtle (Dermochelys coriacea) female, returning to sea after laying eggs in nest on beach, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Adult Common tern (Sterna hirundo) brooding two chicks. Long Island, New York, USA. June
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Black-naped monarch / Black-naped blue flycatcher (Hypothymis azurea) male, with insect prey in beak, feeding three chicks at nest, Wilpattu National Park, Sri Lanka
Great spotted woodpecker (Dendrocopos major) male parent feeding juvenile. Germany. June
Canada goose (Branta canadensis) female, in defensive posture, with brood of goslings on lake, Druridge Pools Nature Reserve, Northumberland, UK. May
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
Red-backed toadlet (Pseudophryne coriacea) male, guarding eggs in nest under rotten log, Border Ranges, Queensland, Australia
Great crested grebe (Podiceps cristatus) carrying a chick on its back in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. May
Female Polar bear (Ursus maritimus) with cub in snow, Churchill, Canada. November
Konik wild horse (Equus ferus caballus) foal suckling its mother, Meinerswijk nature reserve, near Arnhem, the Netherlands. May
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
African wild dog (Lycaon pictus) alpha female suckling her 12 pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Limpkin (Aramus guarauna) female feeding and teaching her chick. Myakka River State Park, Florida, USA
Puma (Puma concolor) female, grooming her cub, Torres del Paine National Park, Magallanes, Chile
Leatherback turtle (Dermochelys coriacea) female, in shallow river, trying to find her way back to the sea after laying eggs on a sandbar, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Common tern (Sterna hirundo) adult in breeding plumage flying, carrying fish for young, Long Island, New York, USA, June
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Female Nursery web spider (Pisaura mirabilis) carrying egg sac, Peak District National Park, Derbyshire, UK. June
Female Leopard (Panthera pardus) carrying cub in her mouth to their new den, Kgalagadi Transfrontier Park, South Africa
Female Kingfisher (Alcedo atthis) covering chicks, aged one day, with her wings in an artificial nest, Italy
Male Kingfisher (Alcedo atthis) sitting on eggs for his brooding period in an artificial nest, Italy
Male Kingfisher (Alcedo atthis) feeding chicks, aged 9 days, with a large fish in artificial nest, Italy
Mother White-tailed deer (Odocoileus virginianus) allowing young to suckle, Finland. September
Adult Swift foxes (Vulpes velox) caring for pup at den, Montana, USA. June
Polar bear (Ursus maritimus) mother and cubs rubbing themselves against mounds of snow, Robeson Channel, North Greenland. August
Southern Tamandua (Tamandua tetradactyla) female carrying young on her back, Northern Pantanal, Mato Grosso State, Brazil, South America
Male Dendtric jawfish (Opistognathus dendriticus) with incubating eggs in his mouth, Dampier Strait, Raja Ampat, Indonesia, Pacific Ocean
Blue-footed booby (Sula nebouxii), incubating eggs with webbed feet, Seymour Island, Galapagos, South America
Swallow-tailed gull (Creagrus furcatus), pair guarding small chick, Genovesa Island, Galapagos, South America
Brown pelican (Pelecanus occidentalis) pair in breeding plumage, collecting nesting material, Santa Cruz Island, Galapagos, South America
Fishing cat (Prionailurus viverrinus) with two kittens, age 4 weeks, learning to hunt for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss
White stork (Ciconia ciconia) flying back to its nest in Oak (Quercus robur) tree to feed its three large chicks, Knepp Estate, Sussex, UK, June
Adult White stork (Ciconia ciconia) standing beside three large chicks on their nest, Knepp Estate, Sussex, UK, June
Panamic sergeant major (Abudefduf troschelii) guarding egg mass, removing seastar that was coming to close to egg mass, Punta Resbaladilla, south of El Tecolote, Gulf of California (Sea of Cortez)
Giant damselfish (Microspathodon dorsalis) guarding egg mass, Coronado Island, Loreto Bay National Park, Gulf of California (Sea of Cortez), Mexico, August
Coyote (Canis latrans) female with newborn pups. Yellowstone National Park, Wyoming, USA
Arboreal tarantula (Lampropelma sp. ) female guarding spiderlings at entrance to hollowArboreal tarantula (Lampropelma sp.) female guarding spiderlings at entrance to hollow retreat, 4 metres up a tree. Danum Valley, Sabah, Borneo
Swift fox (Vulpes velox) female cleans the fur of one of her kits while another tries to nurse, Eastern Colorado, USA, May
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Barbary macaque (Macaca sylvanus) baby sheltering between arms of mother and another adult. Gibraltar Nature Reserve, Gibraltar. August
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar