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Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Fire / European salamander (Salamandra salamandra) Plitvice Lakes National Park, Croatia. November
RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the packground sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia
Underside of Scarlet waxcap fungus (Hygrocybe coccinea), showing stipe and gills. Monmouthshire, Wales, UK, October. Focus-stacked image
Fallen Maple leaves (Acer sp. ) in autumnFallen Maple leaves (Acer sp.) in autumn
Lake Baikal landscape in autumn, trees reflected in water. Zabaikalsky National Park, Siberia, Russia, September 2013
Gills of Honey Waxcap (Hygrocybe coccinea). The National Forest, Leicestershire, UK, October
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Autumn bramble leaves (Rubus fruticosa) Hindringham, North Norfolk, England
Silhouette of Red deer (Cervus elaphus) stag, hind and fawns at sunset, Norfolk, UK
Pattern of green yellow, and red Maple (Acer sp. )leaves in autumn, North Chagrin ReservationPattern of green yellow, and red Maple (Acer sp.)leaves in autumn, North Chagrin Reservation, Ohio, USA, October
Golden Scalycap fungi (Pholiota aurivella) on Beech tree, Sussex, England
European beech (Fagus sylvatica) trees in autumn, Pollino National Park, Basilicata, Italy, November 2008
Silhouette of Red Deer (Cervus elaphus) stag calling during rut, Dyrehaven, Denmark
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Orange jewel anemones (Corynactis viridis) Sula Sgeir, Scotland, UK, February
An Orange lacewing butterfly (Cethosia penthesilea), Cairns Botanical Gardens, Queensland, Australia. Captive. August
Orange coloured Seashore Lichens (Xanthoria parietina) and Sea Ivory (Ramalina siliquosa) UK, August
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Bands of colour created by clouds at dusk, Tartumaa, Estonia, May
Orange-barred sulphur butterfly (Phoebis philea) pe. Costa Rica
Wire coral colonies (Cirrhipathes spiralis), two spirals. Reef in Komodo area, Indonesia
Red ruffed lemur (Varecia rubra) female in understorey of lowland rainforest. Masoala National Park, north-east Madagascar
Cuckoo wrasse {Labrus mixtus / bicamulatus} male, Channel Islands, UK
Marsh fritillary (Euphydryas aurinia) on pink flower. South of Casteil, Pyrenees Orientales, south west France. May
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
RF- Orang utan (Pongo pygmaeus) head portrait of dominant male and first orangutan in world to cataracts operated on with eyesight restored
Nudibranch (Kaloplocamus acutus) Bluff Island, Hong Kong UNESCO Global Geopark, Hong Kong, China
Sea apple, eating, using arm to transfer food into mouth. Ninepin Group or Kwo Chau Islands, Sai Kung District of Hong Kong, China
Sun / Tube coral (Tubastraea sp) with tentacles extended to catch zooplankton. Gulf of Bohai, Yellow Sea. Penglai / Dengzhou / Tengchow, Yantai, Shandong Province, China
Ascidian / Sea squirt (Ascidiacea) close-up. Bohai Sea, Yellow Sea. Zhifu Island, Shandong Province, China
Eucalyptus trees killed by the drought which lasted from 1996-2011, Lake Eucumbene, New South Wales, Australia
Laguna Miluni a reservoir fed by glacial melt water from the Andean peak of Huayna Potosi. As climate change causes the glaciers to melt, the water supply for La Paz
Colourful lakes below the peak of Chacaltaya, lake discoloured by mine effluent. Andes, Bolivia. October 2015
Red slug (Arion rufus) Vosges, France, September
Tassled scorpionfish (Scorpaenopsis oxycephala) swimming just above sea bed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Giant ostracod {Gigantocypris mulleri} from between 718 and 1864m, Mid-Atlantic Ridge, North Atlantic Ocean
Giant ostracod {Gigantocypris mulleri} from between 867 and 1805m, Mid-Atlantic Ridge, North Atlantic Ocean
RF- Rufous Hummingbird (Selasphorus rufus) female feeding in flight on Scarlet Gilia (Ipomopsis aggregata). Gila National Forest, New Mexico, USA. August
Fire lily (Lilium bulbiferrum) flower, Triglav National Park, Slovenia, June 2009