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Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Raccoon (Procyon lotor) approaching with curiousity. Stanley Park, Vancouver, British Columbia, Canada. August
Silky sifaka (Propithecus candidus) female with baby sitting amongst branches in understorey of mid-altitude montane rainforest. Marojejy National Park, north east Madagascar
RF- Stump tailed chameleon (Brookesia superciliaris) portrait, Andasibe-Mantadia National Park, Eastern Madagascar
Canary fish (Epinephelus marginatus) in reef. El Hierro, Canary Islands
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Hawfinch 1+Coccothraustes coccothraustes+2 perched on branch amongst Hawthorn 1+Crataegus sp+2 berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February
Bengal tiger (Panthera tigris), portrait. Bandhavgarh National Park, India. Photo© Phillip Ross/Felis Images
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Gannet (Morus bassanus) chick with fluffy with mouth open. Shetland Islands, Scotland, UK, August
American crocodile (Crocodylus acutus) with mouth open over seagrass bed, Chinchorro Banks Biosphere Reserve, Quintana Roo, Mexico
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Hippopotamus (Hippopotamus amphibius) splashing in pool in evening light. Mana Pools National Park, Zimbabwe
Curious juvenile Grey seal (Halichoerus grypus) Donna Nook, Lincolnshire, England, January
Polar bear (Ursus maritimus) young bear in freezing water during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, Beaufort Sea
Hamerkop (Scopus umbretta) tossing frog prey, Mkuze, South Africa
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Great reed warbler (Arcocephalus arundinaceus) chick begging for food, with bokeh affect from water, Pusztaszer, Hungary, June
Brown Duckling cheeping, against white background
Common / Eurasian crane (Grus grus) calling at sunset, silhouetted. Lake Hornborga, Sweden, April
Hanuman langur / Northern Plains Grey Langur (Semnopithecus entellus) drinking from a water tap at Ranthambore fort. Ranthambore National Park, Rajasthan, India
Razorbill (Alca torda) pair calling, between soft focus vegetation. Great Saltee, Saltee Islands, Co. Wexford, Ireland, June
Whaleshark (Rhincodon typus) swimming and filtering fish eggs from the water. A behaviour known as ram-feeding. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
A portrait of a Mediterranean anthias (Anthias anthias) against open water. Secca Del Papa, Tavolara Marine Protected Area, Sardinia, Italy. Mediterranean Sea
Male Ibex (Capra ibex), Aiguilles Rouges (Red Peaks) National Nature Reserve, Haute-Savoie, France, June
Asiatic black / Moon bear (Ursus thibetanus) head portrait with tongue hanging out, captive
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
Cheetah (Acinonyx jubatus) cub aged 6 months yawning, Masai-Mara Game Reserve, Kenya. Vulnerable species
Amazon river dolphin (Inia geoffrensis), Rio Negro, Amazonia, Brazil
RF- Head portrait of Giant panda (Ailuropoda melanoleuca) chewing on bamboo in snow, captive born in 2000. Occurs in China
Blue Whales (Balaenoptera musculus) throat inflated, filter feeding at sea surface. California, USA. Sequence 1 of 2
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Blue Bearded Collie puppy, 3 months, yawning
West Highland White Terrier sitting
Domestic dog, Border Collie lying on grass, panting
Young Tigress (Panthera tigris), 22 month, being frightened by a dead leaf. Banghavgarh National Park, India, April
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Whale Shark (Rhincodon typus), eating krill and plankton at the surface. Gulf of Mexico, Mexico, North America, August
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)