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Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Amazon river dolphin (Inia geoffrensis), Rio Negro, Amazonia, Brazil
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Keel-billed toucan (Ramphastos sulfuratus) feeding, tossing fruit seed in beak, perched on branch. Boca Tapada, Costa Rica. Sequence 2/2
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Smalltooth sand tiger shark (Odontaspis ferox), view from below. El Hierro. Canary Islands
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Aerial view of Fin whale (Balaenoptera physalus) lunge-feeding, with mouth open and throat pouch distended, southern Sea of Cortez (Gulf of California), Baja California, Mexico
Yorkshire Terrier mother, Evie, with her two Yorkishire terrier x Poodle Yorkipoo pups, age 7 weeks
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
Black spider monkey (Ateles chamek) calling, captive, Peru
Red howler monkey (Alouatta seniculus) howling, captive, digitally modified
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil
Illuminated netdevil (Linophryne arborifera), female specimen, from deepsea Atlantic Ocean near Cape Verde at a depth of 1900m
King cobra (Ophiophagus hannah) in strike pose with mouth open, tongue out and glottis (hole like structure in mouth) clearly visible. Malaysia
Black Cocker Spaniel puppy, against white background
Golden Retriever dog pup, Oscar, 3 months, in play-bow, against white background
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
Leopard (Panthera pardus) female yawning, Samburu game reserve, Kenya
Dalmatian pelican (Pelecanus crispus) low angle perspective of open bill, Lake Kerkini, Greece, February. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Tiger moray (Enchelycore anatina) portrait, Tenerife, Canary Islands
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Fangtooth moray (Enchelycore anatina), close up of eye and open mouth with teeth. Tenerife, Canary Islands
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
