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American quarter horse, three standing in snow. Alberta, Canada. February
Moss-covered Sitka spruce (Picea sitchensis) and sword fern (Polystichum munitum) in the Hoh Rain Forest, Olympic National Park, Washington, USA, June
Frost on soaptree yuccas (Yucca elata) and winter morning fog, in the gypsum sand dunes of White Sands National Monument, New Mexico, USA. January
Flannel bush (Fremontodendron California Glory ) exposed and overflowing nectaries fluorescuing in UV light. Native to southwest USA
Southern magnolia / Bull bay (Magnolia grandiflora) flower. The opening and closing of petals controls the time beetle pollinators enter and leave the flower
Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Southern catalpa (Catalpa bignonioides) flowers. Recently opened with yellow nectar guides, purple in older pollinated flowers. Surrey, England, UK. Native to USA
Buacaille Etive Mor and River Etive in winter, Rannoch Moor, Glencoe, Scotland, UK.December
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
North American porcupine (Erethizon dorsatum) in snow, Vermont, USA. (Habituated rescued individual returned to the wild)
Male and female White-tailed deer (Odocoileus virginianus) with snowman, New York, USA, February
Blinded sphinx moth (Paonias excaecatus) Lac-Drolet Province, Quebec, Canada
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
American alligator (Alligator mississippiensis) emerging from water, in evening light. Myakka River State Park, Florida, USA. March
Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind, in the plankton-rich green waters of La Paz bay
Berkshire piglet in a basket among squashes in early autumn; Rhode Island, USA. October
Beaver pond with beaver lodge (on the left side) and trees reflected in autumn. Acadia National Park, Maine, USA. October
Moose (Alces alces) bull, crossing mountain river after sunset. Grand Teton National Park, Wyoming, USA. October 2017
Autumnal Quaking aspen (Populus tremuloides) and pine tree forest, along the Klondike Highway near Moose Creek, Yukon Territories, Canada, September 2013
Autumnal Quaking aspen tree (Populus tremuloides) at Nares Lake, with Montana Mountain beyond, near Carcross, Yukon Territories, Canada, September 2013
American robin (Turdus migratorius) feeding on Winterberry holly (Ilex sp) berries, New York, USA, November
Pine tree (Pinus sp) in winter, Bryce Canyon National Park, Utah, USA, January
Snowy egret (Egretta thula) shows its breeding plumage. St. Johns Management Area, Florida, USA, March
Western screech owl (Megascops kennicottii) is camouflaged as it looks from a tree cavity. Bosque del Apache National Wildlife Refuge, New Mexico, USA, January
Soaptree yucca (Yucca elata) at sunset. White Sands National Monument, New Mexico, USA. January
Long-finned pilot whale (Globicephala melas) off Nova Scotia, Canada. October
Atlantic bluefin tuna (Thunnus thynnus) off Antigonish, Nova Scotia, Canada. October
Overhead view of a blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
A group of frilled anemones (Metridium senile) off Bonaventure Island in the Gulf of Saint Lawrence, Quebec, Canada. September
Chum salmon (Oncorhynchus keta) migrate up a small river near Bella Bella, British Columbia, Canada. September
Juvenile Pollock (Pollachius virens) school within the protective fronds of kelp near Port Joli, Nova Scotia, Canada. August
Underneath view of a North American beaver (Castor canadensis) swimming under the ice of a frozen river. Quebec, Canada. October
Aerial view of Carters Beach on the Southshore of Nova Scotia, Canada. August
Sea Otter, (Enhydra lutris), young male, Elkhorn Slough, California, USA. January
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July
American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Common loon (Gavia immer) adult in breeding plumage, preening in the last light of day, Michigan, USA, June
Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male perched in marsh vegetation (Phragmites) in foreground. Bear River Migratory Bird Refuge, view toward east
Indigo Bunting (Passerina cyanea) male in breeding plumage perched in Crabapple (Malus sp.) blossom in spring, Ithaca, New York, USA, May
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January