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Spiny dogfish (Squalus acanthias) portrait. Rhode Island, New England, USA. August
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Western hemlock (Tsuga heterophylla) in Douglas-fir forest (Pseudotsuga menziesii), Oregon, USA, June
Common collared lizard (Crotaphytus collaris auriceps) male basking on rock. Arches National Park, Utah, USA. May
North American Beaver (Castor canadensis) grooming. Acadia National Park, Maine, USA
Percheron horses, two including one dappled grey walking through snow, frosty trees in background. Alberta, Canada. February
Arctic wolf in snow (Canis lupus arctos), Minnesota, USA. January. Controlled situation
RF - Mustang mare and foal standing in front of water body. Red Desert Complex, Wyoming. July. (This image may be licensed either as rights managed or royalty free.)
Percheron horse, group running uphill through snow, one dappled grey leading black horses, snow covered forest in background. Alberta, Canada. February
Raccoon (Procyon lotor) female with cub among flowers, Acadia National Park, Maine, USA
Portrait of a wild adult female Bobcat (Lynx rufus) with Hispid cotton rat (Sigmodon hispidus) prey, Texas, USA. September
Portrait of a wild female Bobcat (Lynx rufus) kitten, Texas, USA. September
Portrait of a wild female Bobcat (Lynx rufus) kitten playing, Texas, USA. September
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
Autumn quaking aspen trees (Populus tremuloides) below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Canopy jumping spider (Phidippus otiosus) male, North Florida, USA, September
Great horned owl (Bubo virgininus) with chicks in nest in Saguaro cactus (Carnegiea gigantea) Sonoran desert, Arizona, USA
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Canada Goose (Branta canadensis) in flight. Gilbert Water Ranch, Arizona, USA. January
Bald eagle (Haliaeetus leucocephalus) portrait, captive, occurs in North America
Autumnal boreal forest with lake, Silver Trail, near Mayo, Yukon Territories, Canada, September 2013
Stellers jay (Cyanocitta stelleri) perched on branch. British Columbia, Canada, June
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Osprey (Pandion haliaetus) with Atlantic needlefish (Strongylura marina) it has caught. Blue Cypress Lake, Florida, USA, April
Hybrid Poplar tree plantation in autumn (Populs sp. ). Boardman Tree Farm, near IrrigonHybrid Poplar tree plantation in autumn (Populs sp.). Boardman Tree Farm, near Irrigon, Oregon, USA. October
Blue shark (Prionace glauca) with a pair of pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Sugar kelp (Sacchinaria latissima) off Nova Scotia, Canada. September
A female American black bear (Ursus americanus) and her young cub, Nova Scotia, Canada. July
Invasive Lionfish (Pterois volitans) which have taken over and are wiping out native fish in the Atlantic ocean. The highest densities are in the northern gulf of Mexico. Destin, Florida, USA
North American beaver (Castor canadensis) on dam. Martinez, California, USA. June
Baltimore oriole (Icterus galbula) male perched in pear (Pyrus sp.) blossom, eastern redbud in background, New York, USA. May
American Robin (Turdus migratorius), feeding on winterberry (Ilex) fruits in winter, New York, USA, December
American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April
Clarks grebes (Aechmophorus clarkii) courting pair performing the Weed Ceremony in which they slowly circle breast to breast holding pond weed as nesting material, Lake Hodges, Escondido, California
Cinnamon teal (Anas cyanoptera), male taking flight from water, Orange County, California, USA, April
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Northern cardinal (Cardinalis cardinalis) male perched amid snow-covered branches, New York, USA, February