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Yellow Labrador Retriever standing in glasswort against a blue sky. Monterey Peninsula, California, USA, February
Young Tigress (Panthera tigris), 22 month, being frightened by a dead leaf. Banghavgarh National Park, India, April
Jinshanling Section of the Great Wall of China, a highly authentic part of the wall that has not needed restoration for 400 years. November 2009
Rice-growing fields on hillside terraces, in use since the Yuan dynasty (about 13th century). China, November 2009
Huge flock of Common starlings (Sturnus vulgaris) flying in to roost over the reed beds at dusk, Ham Wall nature reserve, Somerset, UK, January
Portrait of a wild Horse (Equus caballus). The foothills of southwest Reno, Nevada, USA, May
The snow at Horsetail Falls begins to melt in mid-February, enabling the falls to resume once again. The clear sky and low sun angle ignite the rock wall during sundown
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Female Rocky Mountain bighorn sheep ewe (Ovis canadensis canadensis) warning her newborn lamb that a rocky ledge is too close and dangerous
Honey bee (Apis mellifera) approaching a red poppy (Papaver rhoeas) in a small meadow of wildflowers. Nevada, USA, July
Cosmos (Cosmos bipinnatus), Bachelor buttons / Cornflowers (Centaurea cyanus), and Common poppies (Papaver rhoeas) in a small meadow. Nevada, USA, July
Brown bear (Ursus arctos) strolling along the beach in the rain, looking towards the camera. Lake Clark National Park, Alaska, USA, July
Two Brown bears (Ursus arctos) fighting in woodland wetlands, Kuhmo, Finland, July
Beginnings of the Northern lights in night sky in winter with conifer trees laden with snow, Kuusamo, northern Finland, February 2009
Common raven (Corvus corax) perched in snow beside birch tree, Finland, March
Golden eagle (Aquila chrysaetos) about to land on snow, Oulanka NP, Finland, February
A snowy mountain peak lit by low crepuscular light before sunset. Ordesa National Park, Pyrenees, Aragon, Spain, February 2009
Poets Daffodil (Narcissus poeticus). Montseny Nature Reserve, Barcelona province, Catalonia, Spain, May
Whale Shark (Rhincodon typus), eating krill and plankton at the surface. Gulf of Mexico, Mexico, North America, August
Dawn mist on the fields around the village of South Harting. South Downs, Sussex, UK, September 2009
Silver Birch (Betula verrucosa) reflected in Loch Beinn a Mheadhoin. Glen Affric National Nature Reserve, Scotland, December
Panther Chameleon {Furcifer pardalis} climbing along branch on beach, Masoala Peninsula National Park, north east Madagascar
Tree covered with rime ice standing in snow-covered field, Ithaca, New York, USA. January 2010
Fragrant water lilies (Nymphaea odorata) in Lake Skadar, Montenegro, May 2008
Gannet colony at Muriwai, North Island, New Zealand
Southern plains grey / Hanuman langur {Semnopithecus dussumieri} two adolescents sitting, Thar desert, Rajasthan, India
Rainbow in spray of Seljalandsfoss waterfall, Iceland
Cherhill white horse, first cut into chalk downland in 1780, Wiltshire, UK, spring 2009
Lammergier {Gypaetus barbatus} soaring over the Himalayas, northern India
Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Grey Heron (Ardea cinerea) leaving footprints while walking through snow. The Netherlands, January
Mute Swan (Cygnus olor) standing on frosty grass. The Netherlands, March
Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June
Portrait of Grey Heron (Ardea cinerea) looking directly at lens. The Netherlands, June
Konik Wild Horse (Equus ferus caballus) in grasslands with Purple loosestrife flowers (Lythrum salicaria) in foreground. The Netherlands, July
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Reef with Pygmy Sweepers (Parapriacanthus ransonetti), Barrel Sponge (Xestospongia testudinaria), and soft corals. Komodo National Park, Indonesia, October
Close up of feathers of cock Pheasant (Phasianus colchicus)
Apollo butterfly (Parnassius apollo) on thistle head in alpine meadow. Nordtirol, Austrian Alps
Brown Hare (Lepus europaeus) portrait gazing alertly. Perthshire, Scotland, June
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Brown bear (Ursus arctos) walking beside lake, Kamchatka, Far east Russia, August
Alpine meadows with Caucasian scabious (Scabiosa caucasica) in Bezengi gorge, July, 3000m. W-C Caucasus, Kabardino-Balkarskiy nature reserve, Russia
Brown bear (Ursus arctos) standing beside lake looking for salmon, Kamchatka, Far east Russia, August
Wood Ant (Formica rufa) workers on top of their nest synchronise ejection of formic acid droplets to maximise deterrent effect on potential predator, UK
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)
