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Zanzibar red colobus (Piliocolobus kirkii) in tree, Jozany Forest, Zanzibar, Tanzania. Highly endangered
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
White rhinoceros (Ceratotherium simum) mother and calf browsing. Entabeni Safari Conservancy, Limpopo region, Waterberg, South Africa, October
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Male Marsh pride lion (Panthera leo) on a moonless night, Masai Mara, Kenya. Taken with infra red camera, September
Marine Iguana (Amblyrhynchus cristatus) on rocks with wave splash. Galapagos, Ecuador
Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March
Heermanns Gulls (Larus heermanni) in flight. Ensenada, Baja California Peninsula, Mexico, December
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
American Flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
Polar bear (Ursus maritimus) sow and cub sliding on their backs over the pack ice, much like ice skating, 1002 coastal plain of the Arctic National Wildlife Refuge, Alaska, USA
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Long Horn cattle in meadow, Norfolk, UK July
Ash (Fraxinus excelsior) fresh leaf in spring, Dorset, UK April
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Looking out across sea from deserted beach, Christmas Island / Kiritimati, July 2008
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Big fin reef squid (Sepioteuthis lessoniana) descends from open water to the reef at night. Arafur Sea, Triton Bay, West Papua, Indonesia
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Poppies flowering in a field, near Norcia, Umbria, Italy. May 2005
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
Aerial view of patterns created by agricultural work on land, Los Monegros, Zaragoza Province, Aragon, Spain, July 2011
Robin (Erithacus rubecula) perched on rosehip branch during snowfall. UK, December
Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia
A band of semi-feral Quarter mares and foals running across a creek, Estancia Don Amerigo, Chaco, Paraguay, January 2012
Malachite kingfisher (Alcedo cristata) perched on rush, Intaka Island, Cape Town, South Africa, February
Common peafowl (Pavo cristatus) perched on trees at dawn, Western Ghats, Southern India
Cardiff Bay in the evening, with meadow flowers on the Barrage. Cardiff, Wales, July 2011
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Asiatic lionesses and cubs (Panthera leo persica) drinking from pool, Gir Forest NP, Gujarat, India. Water is scarce in the Gir
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Clouded leopard (Neofelis nebulosa) Guwahati, Assam, India, captive
Mistle Thrush (Turdus viscivorus) in snow. Hertfordshire, England, UK, February
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
