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Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Two newly fledged burrowing owl chicks (Athene cunicularia) one being groomed by its mother (far left) Pantanal, Brazil. WINNER
California sea lion (Zalophus californianus) looking down from the surface. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Three Ring-tailed Lemurs (Lemur catta) sun basking at dawn. Berenty Private Reserve, southern Madagascar. (digitally stitched image)
RF- Grape hyacinths (Muscari) and Daffodils (Narcissus sp) in flower, Norfolk, UK, March
Giant Water Lily (Victoria amazonica) flower in a lagoon at Porto Jofre, off the Cuiaba River, northern Pantanal, Mato Grosso State, Brazil, South America
RF- Grey seal (Halichoerus grypus) pups on Norfolk Beach, England, UK, December
RF- European hedgehog (Erinaceus europaeus) orphan, Jarfalla, Sweden. August
Baby Olive Ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the beach where it hatched, Ostional, Costa Rica. October
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
Bengal Tigers (Panthera tigris tigris) adult female drinking water, her juvenile cub beside her. Bandhavgarh National Park, India, April
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, amongst forest vegetation. Endangered. Bandhavgarh National Park, India. Non-ex
Bengal Tiger (Panthera tigris tigris) young cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle July 2008
RF- Siberian tiger (Panthera tigris altaica) young cub, lying down, looking away, captive
RF- Tiger (Panthera tigris) 14-month Lakshmi cub resting in tree, Bandhavgarh National Park, India. Endangered species
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species
RF- Rear view of male Bengal tiger (Panthera tigris tigris) walking on track. Bandhavgarh National Park, India. Endangered species
RF- Holywell Bay in evening light and sand ripples, near Newquay, Cornwall, UK. June 2015
Tufted duck (Aythya fuligula) on water on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. June
RF - Reindeer (Rangifer tarandus) walking up slope in snow. Svalbard, Norway, April. (This image may be licensed either as rights managed or royalty free.)
RF - Arctic fox (Alopex lagopus) in winter pelage, camouflaged in snow. Svalbard, Norway, April (This image may be licensed either as rights managed or royalty free.)
Licancabur volcano and surrounding mountains reflected in waters of a severe flood caused by climate change. Los Flamencos National Reserve, Antofagasta, Chile. February 2019
Kingfisher (Alcedo atthis) perched on branch, Black Forest, Baden-Wurttemberg, Germany
Silver birch (Betula pendula) in autumn fog, Glen Affric National Nature Reserve, Werter Ross, Scotland, UK. October
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Elephant (Loxodonta africana) female and calf under tree in the rain, Masai-Mara game reserve, Kenya
Quiver tree (Aloe dichotoma) with the Milky Way at night, and light pollution from town in the distance, Keetmanshoop, Namibia. Colours accentuated digitally
Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia
RF - Siberian tiger (Panthera tigris altaica) female grooming cub age three months, captive. (This image may be licensed either as rights managed or royalty free.)
RF - Arctic fox (Alopex lagopus) sitting with summer pelage, tuft of fur on head. Svalbard, Norway, July. (This image may be licensed either as rights managed or royalty free.)
Red Fox (Vulpes Vulpes) scratching in sunset light, North London, England, UK, June
Forest lochan at dawn, Abernethy National Nature Reserve, Cairngorms National Park, Scotland, September 2014
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Ancient Beech trees (Fagus sylvatica), Lineover Wood, Gloucestershire UK. The second largest Beech tree in England in the foreground. November 2015
Mountains covered in snow, Chamonix, France, in winter of December 2011
Whale shark (Rhincodon typus) swimming through sun beams. This individual was following blue whales around to feed on their faeces. Indian Ocean, off Sri Lanka
Aquatic plants in water, Donana NP, Andalucia, Spain
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Centuries old English oak / pedunculate oak (Quercus robur) in late summer / autumn, Denmark. September 2018
RF - Banded darter (Sympetrum pedemontanum) group of five resting, Netherlands, September
