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Roseate Tern (Sterna dougallii) in flight in front of building, Madeira Portugal August
American Red Fox (Vulpes vulpes) cubs playing with its mother. Grand Teton National Park, USA, June
American Purple Gallinule (Porphyrio martinica) displaying brightly coloured wings. Everglades National Park, Florida, USA, March
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National Park, Chile
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Common frog (Rana temporaria) leaping into a pond, controlled conditions, UK
Ural Owl (Stix uralensis) perched in snowy tree, Kuusamo Finland February
Wild Cat (Felis silvestris) in mist. Vosges, France, August
Tibetan antelope (Pantholops hodgsoni), two males, Kekexili, Qinghai, Tibetan Plateau, China, December
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
Common poppy (Papaver rhoeas) bud, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
View from the top of the Ozarde waterfalls near Koyna Lake during the monsoon. Western Ghats, India, August 2010
Rainforest interior views, looking up
Magnolia Rustica rubra in flower in garden, UK, May
RF- Great grey owl (Strix nebulosa) in flight, Finland, March
RF- Razorbill (Alca torda) resting on rocks. Great Saltee Island, County Wexford, Ireland. April
Cornflowers (Centaurea cyanus) in fallow fields, near Sugano, Orvieto, Italy, June
Elder flower orchid (Dactylorhiza sambucina) in its two colour forms, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May
White tailed eagle (Haliaeetus albicilla) in flight with snowy background, Rausu, Japan, February
Portrait of male Reindeer (Rangifer tarandus) with large antlers, in velvet. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June
Eastern Island showing the WW2 airstrip which was used to launch the initial attack on the Japanese fleet for the Battle of Midway, Midway, Central Pacific
Alpine Snowbell (Soldanella alpina) in flower, above Madoona di Campiglio, Brenta Dolomites, Italy, June 2010
RF - Delicate grass and flowers silhouetted black against white background
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
RF- Eurasian Eagle owl (Bubo bubo) adult perched on rocky outcrop with the Super Full Moon, September 28th 2015, Southern Norway. August. Multiple exposure
Snowy owl (Bubo scandiaca) close up of female landing on snow covered ground, Canada, February. Magic Moments book plate
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Stellers Sea Eagles (Haliaeetus pelagicus) fighting on ice floe, Hokkaido, Japan. February
Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014
Buff-tailed Bumblebee (Bombus terrestris) worker feeding on Ox-eye Daisy (Leucanthemum vulgare). Peak District National Park, Derbyshire, UK. July
RF - Red Squirrel (Sciurus vulgaris) adult in summer coat leaping onto fallen log. Scotland, UK. September
Marbled white butterfly (Melanargia galathea) roosting on grasses, England, UK, July
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Red Squirrel (Sciurus vulgaris) adult sitting and feeding in snowfall. Cairngorms National Park, Scotland, UK, March
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
RF- Serotine bat (Vespertilio serotinus) in flight above Hogweed. Captive, UK
RF- Red squirrel in autumn (Sciurus vulgaris). Strathspey, Scotland, UK
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Pasque flowers (Pulsatilla vulgaris) in rain, Lorraine, France, April. 3rd Prize in the Other Nature Topics category of Melvita Nature Images Awards 2013
RF- Sunset on Kimmeridge Bay, Isle of Purbeck, Jurassic Coast World Heritage Site, Dorset, England, UK. November 2008