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Cormorants (Phalacrocorax carbo) in early morning mist, Oisterwijkse Bossen en Vennen nature reserve, Netherlands, November
Great crested grebe (Podiceps cristatus) young playing with a feather next to one of the parents. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. May
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Chinese goral (Naemorhedus griseus) in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Honey bee (Apis mellifera) covered in pollen emerging from nectaring in Crocus (Crocus sp). On nature reserve, Surrey, England, UK. March
Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April
Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
China nawab (Polyura narcaeus) butterfly drinking at thermal seep. Gaoligongshan National Nature Reserve, Yunnan Province, China
China glory flower (Clerodendrum bungei) fruits above red calyxes. Eaten and spread by birds. Tangjiahe National Nature Reserve, Sichuan Province, China
Flowers and leaves on forest floor. Xishuangbanna National Nature Reserve, Yunnan, China
Sancha River flowing through tropical rainforest in dry season. Wild Elephant Valley, Xishuangbanna National Nature Reserve, Yunnan Province, China. April 2007
Dead leaf / Orange oak leaf (Kallima inachus) butterflies feeding on sap on tree trunk. Mimic leaves with wings closed. Wild Elephant Valley, Xishuangbanna, Yunnan Province, China
Marsh fritillary butterfly (Euphydryas aurinia) feeding on Wild garlic / Ramsons (Allium ursinum) at Strawberry Banks, Gloucestershire Wildlife Trust (GWT), Nature Reserve, Gloucestershire, UK. May
Landscape of a fen at dawn withCanada goose (Branta canadensis). Oisterwijkse Bossen en Vennen Nature reserve, Oisterwijk, The Netherlands. April
Cousine Island, a privately owned island and nature reserve, one of the few remaining rat-free islands of the inner Seychelles, off the south-west coast of Praslin Island, seen from Cousin Island
Trees in Hautes Fagnes Nature Reserve in winter after snowfall. This area - Noir Flohay was burnt a few years ago and some trees are still blackened, Hautes Fagnes, Belgian Ardennes, Belgium
RF - Marbled White butterfly (Melanargia galathea) resting on dew covered grasses, Dunsdon Nature Reserve, Devon, UK. July
RF - Mallard (Anas platyrhynchos) female swimming with with young chicks following, Valkenhorst Nature Reserve, The Netherlands June
RF- Two Small red damselflies (Ceriagrion tenellum) covered in morning dew. Arne (RSPB) Nature Reserve, Dorset, UK. August
Silver birch (Betula pendula) woodland in early autumn, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October 2014
Silver birch (Betula pendula) woodland in autumn, Craigellachie National Nature Reserve, Cairngorms National Park, Scotland, UK, October 2014
Scots pine (Pinus sylvestris) trees in Caledonian pine forest at sunrise, Abernethy National Nature Reserve, Cairngorms National Park, Scotland, UK, September 2014
Atlantic puffin (Fratercula arctica) diving, North Rona, Outer Hebrides / Western Isles, Scotland, UK, July
Aerial view over Insh Marshes National Nature Reserve, Cairngorms National Park, Scotland, UK, May 2016
Northern gannet (Morus bassana) in flight, Hermaness National Nature Reserve, Shetland, Scotland, July
RSPB Nature reserve at Titchwell Marsh Norfolk, May
Male Ibex (Capra ibex), Aiguilles Rouges (Red Peaks) National Nature Reserve, Haute-Savoie, France, June
Large male Reindeer (Rangifer tarandus) standing in shallow waters of the Kronotsky lagoon, away from biting insects. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) jumping across stream. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Ranger station in winter with Kamchatka Brown Bear (Ursus arctos beringianus), Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June. Sequence 3 of 3
Kamchatka Brown Bear (Ursus arctos beringianus) on the Pacific coast of Kamchatka. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Ponta de Sao Lourenco Nature Reserve, Madeira
Huge flock of Common starlings (Sturnus vulgaris) flying in to roost over the reed beds at dusk, Ham Wall nature reserve, Somerset, UK, January
Poets Daffodil (Narcissus poeticus). Montseny Nature Reserve, Barcelona province, Catalonia, Spain, May
Silver Birch (Betula verrucosa) reflected in Loch Beinn a Mheadhoin. Glen Affric National Nature Reserve, Scotland, December
Alpine meadows with Caucasian scabious (Scabiosa caucasica) in Bezengi gorge, July, 3000m. W-C Caucasus, Kabardino-Balkarskiy nature reserve, Russia
Alder trees (Alnus sp) reflected in standing water in nature reserve, Berlin, Germany, May 2009
