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Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Manta ray (Mobula alfredi) feeding on plankton aggregated in the lights from a boat at night, North Ari Atoll, Maldives, Indian Ocean
Red grouse (Lagopus lagopus) coming in to land on flowering Heather (Calluna vulgaris), Yorkshire, UK. September
Male Cheetah (Acinonyx jubatus) stretching and yawning on a fallen tree trunk, Okavango Delta, Botswana, Africa
Female Ostrich (Struthio camelus) feeding on desert plants, Kgalagadi Transfrontier Park, South Africa
Japanese pinecone fish (Monocentris japonica) under a ledge in the reef alongside a Giant moray eel (Gymnothorax javanicus) and Banded coral shrimps (Stenopus hispidus), Richelieu Rock
Male Beautiful sunbird (Cinnyris Pulchellus) courtship display, Allahein River, The Gambia
Tiger shark (Galeocerdo cuvier) swimming close to surface at night, Tiger Beach, Bahamas, Caribbean Sea
Great hammerhead shark (Sphyrna mokarran), critically endangered, swimming over sandy seabed accompanied by Blue runner jacks (Caranx crysos) and Remoras (Echeneis sp.), Bimini, Bahamas, Caribbean Sea
Three male Ostriches (Struthio camelus) running and flapping wings in aggressive display, Kgalagadi Transfrontier Park, South Africa
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
RF - Giant moray (Gymnothorax javanicus) with mouth open, looming out of a crevice in coral reef at night, Sharm El Sheikh, Egypt, Red Sea
RF - Male Great white shark (Carcharodon carcharias) swimming beneath the surface, Guadalupe Island, Baja California, Mexico, Pacific Ocean
Manta ray (Mobula alfredi) with mouth open, feeding on plankton near the surface, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean
Wolf (Canis lupus) walking in snow, Yellowstone National Park, USA. January
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Whale shark (Rhincodon typus), an endangered species, feeding on zooplankton in nutrient-rich murky water, being followed by a school of Cobia (Rachycentron canadum), Adang-Rawi Archipelago, Satun
Northern fulmar (Fulmarus glacialis) portrait, Fair Isle, Shetland, Scotland, UK. May
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic, volumetric lighting, Hungary, January
Female Abyssinian ground hornbill (Bucorvus Abyssinicus) walking along riverbank, Allahein River, The Gambia
Wood pigeon (Columba palumbus) perched in a Beech tree (Fagus sylvatica) feeding on buds, Wiltshire, UK, April
Asiatic lion (Panthera leo persica) cub, Gir Forest National Park, Gujarat, India. May
Robin (Erithacus rubecula) perching on a branch, singing, Bavaria, Germany. May
Whale shark (Rhincodon typus) feeding close to fishermens boats in the early morning, Triton Bay, West Papua, Indonesia, Pacific Ocean
Reef manta ray (Mobula alfredi) feeding, with gills and cephalic lobes visible, Dampier strait, Raja Ampat, West Papua Indonesia, Pacific Ocean
Female Eclectus parrot (Eclectus roratus) feeding in rainforest, Cape York, Queensland, Australia
Galapagos land iguana (Conolophus subcristatus) recently reintroduced to Santiago island where it had become extinct, feeding on Yellow cordia (Cordia lutea) leaves, Santiago Island, Galapagos
Two Arctic foxes (Vulpes lagopus), blue-morph with winter coats, playing, Hornstrandir Nature Reserve, Iceland. March
Male Jaguar (Panthera onca) walking through rainforest, Belize, Central America, 2017. Captive. Filmed for the BBC series Big Cats
Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
