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Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Eurasian lynx (Lynx lynx) yawning. Captive
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Giant panda (Ailuropoda melanoleuca) sitting with Bamboo in background. Shenshuping Panda Base, Wolong Nature Reserve, Wenchuan, Sichuan Province, China. Captive
Yorkshire Terrier mother, Evie, with her two Yorkishire terrier x Poodle Yorkipoo pups, age 7 weeks
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Sperm whale (Physeter macrocephalus) at surface, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May
King cobra (Ophiophagus hannah) in strike pose with mouth open, tongue out and glottis (hole like structure in mouth) clearly visible. Malaysia
RF- European brown bear (Ursus arctos) with mouth open, Kuhmo, Finland. July
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Basking shark (Cetorhinus maximus) feeding on plankton in surface waters close to the island of Coll, Inner Hebrides. Scotland, UK. Atlantic Ocean. June
Spotted porcupinefish (Diodon hystrix), inflated with seawater, a defensive behaviour, Hawaii, Pacific Ocean
Frill-necked lizard (Chlamydosaurus kingii) threat display with neck frill extended, Cooktown, Queensland, Australia
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Two Lionesses (Panthera leo) in the rain, one shaking water off and the other yawning. Masai-Mara game reserve, Kenya
Caracal (Caracal caracal) male, standing on rock against blue sky, Spain. Captive, occurs in Africa and Asia
Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp)Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp.), Brooks Falls, Katmai National Park, Alaska. July
Grizzly bear (Ursus arctos) female with cub seeking attention from its mother, Lake Clark National Park, Silver Salmon, Alaska. July
Yacare caiman (Caiman yacare) with mouth open, head portrait, Cuiaba River, Pantanal, Mato Grosso, Brazil
Bluethroat (Luscinia svecica cyanecula) male, perched on tree stump singing, Germany. March
Keel-billed toucan (Ramphastos sulfuratus) perched in Trumpet tree (Cecropia peltata) feeding on fruit, Caribbean slope of Guatemala
Group of California sea lions (Zalophus californianus) playing near the sea surface, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Golden retriever, female, lying down on coastal rocks near ocean, Pemaquid Point, Maine, USA. June
German shepherd dog, female, standing on rocks, portrait, Jacksonville, Oregon, USA. July
Cockapoo puppy, aged 18 weeks, running, portrait
Ragdoll-cross kitten, aged 3 weeks, meowing, portrait
Border collie x Bearded collie puppy, aged 4 months, sitting looking up with tongue out, portrait
Galapagos sea lion (Zalophus wollebaeki) juvenile, jumping and playing in ocean, San Cristobal Island, Galapagos National Park, Pacific Ocean. Endangered
Black-naped monarch / Black-naped blue flycatcher (Hypothymis azurea) male, with insect prey in beak, feeding three chicks at nest, Wilpattu National Park, Sri Lanka
Galapagos fur seal (Arctocephalus galapagoensis) with mouth open, head portrait, Santiago Island, Galapagos National Park, Galapagos Islands. Endangered
RF - Puffin (Fratercula arctica) standing on rock, yawning, Runde, Norway. June. (This image may be licensed either as rights managed or royalty free.)
Reef manta ray (Mobula alfredi) feeding on plankton aggregating in the lights from a boat at night, Ari Atoll, Maldives, Indian Ocean
Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp)Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp.) on a reef, Ari Atoll, Maldives, Indian Ocean
Roundhead parrotfish (Chlorurus strongycephalus) feeding on coral and algae growing on a reef, South Male Atoll, Maldives, Indian Ocean
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Stork-billed kingfisher (Pelargopsis capensis) perched on branch with beak open, Bardia National Park, Terai, Nepal
Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp)Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp.) tree in spring, Ithaca, New York, USA. May
Two Speckled moray eels (Gymnothorax dovii) feeding on a Sea turtle carcass on the seabed, Wolf Island, Galapagos Islands, Pacific Ocean
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador