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Jaguar (Panthera onca) cub nuzzling its mother, Cuaiba River, Pantanal wetlands, Mato Grosso, Brazil
Jaguar (Panthera onca) cub licking its mother, Cuaiba River, Pantanal wetlands, Mato Grosso, Brazil
Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, grazing, being pestered by kid, aged two months, begging for milk, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), alert female with kid aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May
South Andean deer (Hippocamelus bisulcus), adult female with fawn, resting on mountainside. Cerro Castillo National Park, Patagonia, Chile. April
Lions (Panthera leo), female with cub, drinking at waterhole. Etosha National Park, Namibia. August
African elephant (Loxodonta africana), female and calf, crossing plain at sunset. Amboseli National Park, Kenya. July
Gemsbok (Oryx gazella) female and calf walking past sand dune, Sossusvlei UNESCO World Heritage Site, Namib desert, Namib-Naukluft National Park, Namibia
Sika deer (Cervus nippon) doe with fawn standing on rocky outcrop overlooking forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Grey seal (Halichoerus grypus) pup and mother sniffing and nuzzling each other. Donna Nook, Lincolnshire, UK. November
Sulawesi black macaque / Celebes crested macaque (Macaca nigra) troop in forest, female with infant, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Puma (Puma concolor) female, with two cubs, aged six months, sitting on on rocky outcrop overlooking Lake Sarmiento, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Peregrine falcon (Falco peregrinus) female, flying towards nest box with three chicks, aged 18 days, inside, Sagrada Familia Basilica, Catalonia, Spain. May
American oystercatcher (Haematopus palliatus) female with chick resting on beach, Santiago Island, Galapagos National Park, Galapagos Islands
Puma (Puma concolor) female with two cubs, Torres del Paine National Park, Magallanes, Chile
Puma (Puma concolor) female with cub, resting, Torres del Paine National Park, Magallanes, Chile
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Humpback whale female calf (Megaptera novaeangliae) posing with her pectoral fins held together, with her mother resting in the background. Vava'u, Kingdom of Tonga. Pacific Ocean
Milne-Edward's sifaka (Propithecus edwardsi) female with infant sitting in tree, Ranomafana National Park, south east Madagascar. Endangered
Wood duck (Aix sponsa) female, leading brood of ducklings on pond, Acadia National Park, Maine, USA. June
Lioness (Panthera leo) with cub drinking at water hole, Etosha National Park, Namibia
Sperm whale (Physeter macrocephalus) mother and calf, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit. January
Canada goose (Branta canadensis) female, in defensive posture, with brood of goslings on lake, Druridge Pools Nature Reserve, Northumberland, UK. May
Graylag gosling (Anser anser) resting under parent's wing, Pusztaszer, Kiskunsagi National Park, Pusztaszer, Hungary
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
New Forest donkeys, female with foal, standing on grassland, touching noses at dusk, New Forest National Park, Hampshire, England, UK. May
Galapagos sea lion (Zalophus wollebaeki) female with pup lying on rocks, Espanola Island, Galapagos, South America
Rock hyrax / dassie (Procavia capensis), with babies, De Hoop Nature Reserve, Western Cape, South Africa
Female Polar bear (Ursus maritimus) with cub in snow, Churchill, Canada. November
Konik wild horse (Equus ferus caballus) foal suckling its mother, Meinerswijk nature reserve, near Arnhem, the Netherlands. May
Female Gentoo penguin (Pygoscelis papua) with chick on the nest, Port Lockroy, Goudier Island, Antarctic Peninsula, Antarctica
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
African wild dog (Lycaon pictus) alpha female suckling her 12 pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Limpkin (Aramus guarauna) female feeding and teaching her chick. Myakka River State Park, Florida, USA
Puma (Puma concolor) female, grooming her cub, Torres del Paine National Park, Magallanes, Chile
Southern Right whale (Eubalaena australis) mother blowing / spouting at surface, with calf breaching. Peninsula Valdes, Chubut Province, Patagonia, Argentina. August
Aerial view of tourists watching Southern right whales (Eubalaena australis) from the coast, Peninsula Valdes, El Doradillo Nature Reserve, Chubut Province, Patagonia, Argentina. August
Puma (Puma concolor) female with two cubs, aged six months, sitting in fresh snow, Torres del Paine National Park, Patagonia, Chile
Female Galapagos sea lion (Zalophus wollebaeki) with yearling pup resting on beach, Cerro Brujo, San Cristobal Island, Galapagos Islands, Ecuador. Pacific Ocean
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Female Bison (Bison bison) with young calf, walking over snow in front of frost-covered tree, Yellowstone National Park, USA. February
Female Leopard (Panthera pardus) carrying cub in her mouth to their new den, Kgalagadi Transfrontier Park, South Africa
Female Cheetah (Acinonyx jubatus) with juveniles crossing grassland in rain storm, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Gray whale (Eschrichtius robustus), mother and calf, swimming at surface, Magdalena Bay, Baja California, Mexico, Pacific Ocean
Roe deer (Capreolus capreolus) doe with fawn standing in hedgerow, Fife, Scotland, UK. September