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Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Bealach na Ba Pass, Wester Ross, Scotland, UK, November 2015
Rock formations at Brimham Rocks created by variable erosion of soft and hard layers of carboniferous age millstone grit, Harrogate, Yorkshire, England, UK, October 2016
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
View towards Glastonbury Tor over reedbeds at dawn, Somerset, England
Fly agaric (Amanita muscaria) in foggy forest, Los Alcornocales Natural Park, southern Spain, January
Aerial view of Lago Preto, Amazon Rainforest and lake, Peru
Lioness (Panthera leo) sitting during misty sunrise, Okavango Delta, Botswana
Whooper Swan (Cygnus cygnus) male on moorland loch, Isle of Skye, Hebrides, Scotland, December
Hawksbill turtle (Eretmochelys imbricata) pair circling each other above coral reef. Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Great egret (Ardea alba) in snow, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. March
Common cottongrass (Eriophorum angustifolium) flowering, at sunrise, Island Runde, Norway, July
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
RF- Large red damselfly (Pyrrhosoma nymphula) emerging from nymphal case. Cornwall, UK. April
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Scots pine (Pinus sylvestris) forest in evening light, Abernethy, Cairngorms National Park, Scotland, UK, June
Scots pine (Pinus sylvestris) silhoutte in front of Loch Maree and Slioch, Wester Ross, Scotland, UK, November 2014
Sea stacks in morning light, Duncansby Head, John O Groats, Caithness, Scotland, UK, April 2015
View from Isle of Skye across Sound of Rsay to Rona and Torridon Hills, at dawn, Inner Hebrides, Scotland, UK, April 2014
Reflections in Loch Baa at dawn, Rannoch Moor, Glencoe, Lochaber, Scotland, UK, October 2014
Red throated diver (Gavia stellata) on lochan in dawn mist, Cairngorms National Park, Scotland, UK, May
Autumn colours and stormy clouds reflected in the calm waters of Loch Insh, Cairngorms National Park, Scotland, UK, November 2015
Iceberg in the sea outside the Kangia Ilulissat Icefjord. Iceberg from the Sermeg Kujalleg Glacier, UNESCO World Heritage Site, Kalaallit Nunaat, Greenland, August 2014
Sheep (Ovis aries) resting on the Quiraing mountains with low cloud cover, Isle of Skye, Inner Hebrides, Scotland, UK. July
The Pont St-Benezet, Palais des Papes and Rhone River at dusk, Avignon, Provence, France, October 2012
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National Park, Chile
Monochrome image of a Hawthorn tree (Crataegus monogyna) and granite outcrop near Saddle Tor, Dartmoor National Park, Devon, England, UK, January 2011
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
Black-throated diver (Gavia arctica), Finland, June
Aerial of Kossol Passage, Kossol Reef, Palau Islands, North Pacific Ocean
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014
RF- Daffodils (Narcissus) in flower photographed using soft focus technique
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
Loch Beinn a Meadhoin in the mist, Inverness, Scotland, December 2013
Dawn over the mountains and Nam Song River at Vang Vieng, Laos, March 2009
Cheetah (Acinonyx jubatus) female standing below dark storm clouds, Masai-Mara game reserve, Kenya
Three-banded plover (Charadrius tricollaris) in water, with Lion (Panthera leo) sleeping nearby, Ndutu, Tanzania
Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015
Tranquil landscape with African elephant (Loxodonta africana) and rays of sunlight at sunrise, Masai Mara, Kenya
Mute swan (Cygnus olor) pair on misty lake, Amsterdamse Waterleidingduinen Nature Reserve, near Zandvoort, The Netherlands, February
Kirkjufell mountain at dawn with ice in foreground, Snaefellsnes peninsula, Iceland, January 2014