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Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt
Shining honeycreeper (Cyanerpes lucidus) male and Red-legged honeycreeper (Cyanerpes cyaneus) perched on branch with epiphyte. Boca Tapada, Costa Rica
King Penguins (Aptenodytes patagonicus) approached by an Antarctic Fur Seal (Arctocephalus gazella). A few pecks from the penguins caused the seal to retreat
Moths (Lepidoptera) swarming with flight trails, attracted by light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands
Moths (Lepidoptera) with flight trails, attracted to light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands. June
Diverse hard coral growth (Pocillopora damicornis, Pocillopora verrucosa and Millepora dichotoma), flourishes beneath the barren desert cliffs in the Red Sea. Ras Katy, Sinai, Egypt
Dense flock of Common teal (Anas crecca) with a few Wigeon (Anas penelope) and Northern shoveler (Anas clypeata) in flight over others resting on partly frozen flooded marshes
Black-and-white sea flea (Chromopleustes oculatus) on a Sunflower star (Pycnopodia helianthoides), Port Neville, Johnstone Strait, British Columbia, Canada. March
Proliferating anemones (Epiactis prolifera) and Kelp lace bryozoans (Membranipora serrilamella) on Bull kelp (Nereocystis luetkeana), Browning Pass, Queen Charlotte Strait, British Columbia, Canada
Split level shot of kelp and other seaweeds partly exposed at low tide, with coniferous forest in the background. Seen in this photograph are the following
Helmet crab (Telmessus cheiragonus) and Hooded nudibranchs (Melibe leonina), Nigei Island, Queen Charlotte Strait, British Columbia, Canada. October
Coral heads surrounded by Blacktip cardinalfish (Apogon atradorsatus)and juvenile salemas (Xenocys jessiae), Rabida Island, Galapagos
Aerial view of Galapagos sea lion group (Zalophus wollebaeki) swimming with Brown pelicans (Pelecanus occidentalis) followng them to hunt and scavenge
Galapagos lava lizard (Microlophus albemarlensis) on head of Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapagos
Espanola lava lizard (Microlophus delanonis) female with Waved albatross (Phoebastria irrorata) behind. Punta Suarez, Espanola Island, Galapagos
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos
Golden Rays (Rhinoptera steindachneri) swimming with Green turtle (Chenolia mydas). Isabela Island. Galapagos
Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos
Epanola saddelback tortoise (Chelonoidis hoodensis), with Galapagos mockingbird (Mimus parvulus) perched on carapace, Espnola Island, Galapagos
Galapagos fur sea (Arctocephalus galapagoensis) pup, curiously approaching group of Marine iguanas (Amblyrhynchus cristatus) Cape Hammond, Fernandina Island, Galapagos
Wildlife on the coast of Cape Hammond including Galapagos flightless cormorant (Phalacrocorax harrisi), Sally lightfoot crab (Grapsus grapsus) and Marine iguana (Amblyrhynchus cristatus)
Flightless cormorant (Phalacrocorax harrisi) coming ashore amongst crabs. Clearwater Bay, Isabela Island, Galapagos. Sequence 3 of 3
Waved Albatross (Phoebastria irrorata) and Blue-footed Booby (Sula nebouxii) squabbling over nesting sites. Galapagos Islands, Ecuador, June
Schoolmaster snapper (Lutjanus apodus) and Bluestriped grunt (Haemulon sciurus) seeking protection in Elkhorn Coral (Acropora palmata), Jardines de la Reina / Gardens of the Queen National Park
Black grouper (Mycteroperca bonaci), and Caribbean Reef Shark (Carcharhinus perezi), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
African buffalo (Syncerus caffer) herd showing defensive behaviour as an African lioness (Panthera leo) walks by, Mana Pools National Park, Zimbabwe. September
Crinoid (Crinoidea) on Soft coral (Alcyonacea). Derawan Islands, East Kalimantan, Indonesia
Silky shark (Carcharhinus falciformis) close to the surface and near boat with Black jacks / Trevally (Caranx lugubris) and diver in the background, Revillagigedo islands, Mexico. Pacific Ocean
Bramble leaves (Rubus fruticosus) and bracken fronds changing colour in autumn
Kaleidoscopic pattern of Virginia creeper (Parthenocissus quinquefolia) in autumn on a white background
Kaleidoscopic image of Bramble leaves (Rubus fruticosus) and bracken fronds changing colour in autumn, on white background
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
African elephant (Loxodonta africana) two silhouetted at sunset with goose flying overhead, Mkuze, South Africa. Highly commended in the African Wildlife category of the Nature's Best Photography
Great spotted woodpecker (Dendrocopos major) and Grey headed woodpecker (Picus canus) on tree trunk, Germany. December
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Meadow flowers, Fleabane (Erigeron sp) Thistle, Bearded bellfower (Campanula barbata) Common spotted orchid (Dactylorhiza)
Great skua (Stercocarius skua) in flight over Gannet (Morus bassanus) colony, looking for small chicks, whilst Gannets call defensively, Shetland Isles, July
Flamingo tongue (Cyphoma gibbosum) feeding on a seafan (Gorgonia ventalina) Cyphomas feed on corals and concentrate the toxic chemicals into their mantle
Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean
California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
Reef under the surface of shallow waters, at sunset, covered with hard corals, Brush Coral (Acropora hyacinthus) Robust Acropora (Acropora robusta) and other Acropora, Maldives, Indian Ocean
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Two ginger kittens rolling playfully on their sides
Ginger kitten, 7 weeks, sitting between two young Lionhead-Lop rabbits
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
