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Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Three-banded plover (Charadrius tricollaris) in water, with Lion (Panthera leo) sleeping nearby, Ndutu, Tanzania
Tench (Tinca tinca) on the surface of a turbid marsh pond, with Rudd (Scardinius erythrophthalmus) above, Fribourg, Switzerland, 2004
Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
Bobtail / Old English Sheepdog, sitting alongside a longhaired Chihuahua, tan and white coated
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
Cornfield annual summer wildflowers growing on one of the plant charity Landlife sCornfield annual summer wildflowers growing on one of the plant charity Landlifes National Wildflower Farms, St Helens, Merseyside, UK, June
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Brooks urchin shrimp (Allopontonia brooki), commensal among the venomous spines of Variable sea urchin (Asthenosoma varium). On sea floor, Komodo, Indonesia
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Bumblebee (Bombus sp) and Honey bee (Apis mellifera) nectaring on Love-in-a-mist (Nigella damascena) flower. Anthers transferring pollen onto bee backs
Hammerkop / Hammerhead stork (Scopus umbretta) eating a toad, with an African buffalo / Cape buffalo (Syncerus caffer) in the background at night, Zimanga Private Nature Reserve, KwaZulu Natal
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Donkey and horse on the Norfolk Broads with windmill at dawn, Norfolk, UK, October 2008
Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp)Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos
Brown bear Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August
Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp). Anilao, Philippines
Blue mussels (Mytilus edulis) with encrusting marine life, Sula Sgeir, Scotland. UK, July
Elegant anemone (Sagartia elegans) and Plumose anemone (Metridium senile) with fish swimming in plankton bloom in background, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Black-and-white kitten, Solo, 6 weeks, rubbing against F1b toy Goldendoodle (Golden Labrador cross Toy poodle) puppy
African hunting dogs (Lycaon pictus) pack mobbing a Spotted hyena (Crocuta crocuta). Hyenas follow the dogs and attempt to steal their kills, so the dogs harass them whenever they get a chance
Great egret (Ardea alba) at twilight, Lake Csaj, Kiskunsagi National Park, Hungary. July
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica
Green turtle (Chelonia mydas) swimming with snorkelers in the background, Canary Islands, Spain, July
White-throated kingfisher (Halcyon smyrnensis) with Black drongo (Dicrurus macrocercus) Ranthambore National Park, Rajasthan, India
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Katydid (Tettigoniidae) with Giraffe (Giraffa camelopardalis) in the background, Masai Mara National Park, Kenya
Gorgonian sea fans (Melithaea sp. ) with a large school of Pygmy sweepers (ParapriacanthusGorgonian sea fans (Melithaea sp.) with a large school of Pygmy sweepers (Parapriacanthus ransonetti) Andaman Sea, Thailand
Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Sponge crab (Inachus sp) hiding in anemone, Channel Islands, UK June
Soft Coral (Dendronephthya sp) with Blueline snappers (Lutjanus kasmira). Rinca, Komodo National Park, Indonesia
Hoopoe (Upupa epops) with European roller (Coracias garrulus) in the background. Pusztaszer, Hungary. May
Southern elephant seal (Mirounga leonina), male at sunrise with King penguin (Aptenodytes patagonicus) colony in background. St Andrews Bay, South Georgia. October 2017
Chameleon goby (Tridentiger trigonocephalus) with Blue bat star (Patiria pectinifera) in background. Bohai Sea, Yellow Sea. Zhifu Island, Shandong Province, China
Cheetah (Acinonyx jubatus) walking near wildebeest, Masai Mara Nature Reserve, Kenya
Wood anemone (Anemone nemorosa) and Corydalis flowers, (Corydalis solida), Staatsbossen, Texel, the Netherlands, April
Sambar deer (Cervus unicolor), Spotted deer (Axis axis), Wild boar (Sus scrofa) and various birds feeding, with Rajbagh Palace in the background, Ranthambhore National Park, India. April 2010
Peacock (Pavo cristatus) on rock with Palm Squirrel (Funambulus sp) India
Two Lionesses (Panthera leo) trying to defend kill from large pack of Spotted hyaenas (Crocuta crocuta) Masai-Mara game reserve, Kenya
Dingo (Canis lupus dingo) with Silver Gull (Larus navaehollandiae) on beach, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Brown pelicans (Pelecanus occidentalis) on water, split level view, Galapagos
Small heath butterfly (Coenonympha pamphilus) and snail, Ottange, Lorraine, France. August
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya