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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Poppies (Papaver rhoeas) and cornflowers (Centaurea cyanus) near Castellucio di Norcia, Umbria, Italy, July
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
Tabby kitten, Picasso, 9 weeks, head to head with Yellow labrador puppy, 8 weeks
Black and white Border collie bitch with black and white tuxedo kitten age 10 weeks
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Grizzly bear (Ursus arctos horribilis) and Grey wolf (Canis lupus) on beach, Katmai National Park, Alaska, USA, August
Bream (Abramis brama) and European chub (Leuciscus / Squalius cephalus) shoal in a calm part of the Rhine, Schaffhausen, Switzerland, February 2009 WWE OUTDOOR EXHIBITION
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Red-headed woodpeckers (Melanerpes erythrocephalus), pair fighting with European Starling (Sturnus vulgaris) that is trying to take over the woodpeckers nest hole, Montezuma National Wildlife Refuge
Sleepy ginger kitten and lionhead cross rabbit
Peruvian Tarantula (Pamphobeteus sp. ) adult, walking over Humming Frog (ChiasmocleisPeruvian Tarantula (Pamphobeteus sp.) adult, walking over Humming Frog (Chiasmocleis royi) without preying on it. Los Amigos Biological Station, Madre de Dios, Amazonia, Peru
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Various species of moth from tropical rainforest, Danum Valley, Sabah, Borneo. Digital composite
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Anemone prawn (Periclimenes sagittifer) on Snakelocks anemone (Anemonia viridis) Jersey, British Channel Islands, June
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Baby rabbit with a Roborovski Hamster (Phodopus roborovskii) sitting on its head
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
Polar bear (Ursus maritimus) hunting Seal on sea ice. Svalbard, Norway, July 2018
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August
Brooks urchin shrimp (Allopontonia brooki), commensal among the venomous spines of Variable sea urchin (Asthenosoma varium). On sea floor, Komodo, Indonesia
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Butterflies and day flying moths feed from Loofah flower (Luffa cylindrica) and pick up pollen on their legs, underside of thorax and antennae, South Sichuan Bamboo Sea, China
Austral blackbirds (Curaeus curaeus) and Austral thrush (Turdus falcklandii) nectaring on Blue puya (Puya berteroniana). Pollen transferred to birds heads during feeding. Parque Ingles, Chile
Scalloped hammerhead shark (Sphyrna lewini) swims through a school of Pacific creolefish (Paranthias colonus). Wolf Island, Galapagos National Park, Galapagos Islands.. East Pacific Ocean
Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Several mimic goatfish (Mulliodichthys mimicus) hide within a school of Bluestripe snapper (Lutjanus kasmira) on a coral reef. North Ari Atoll, Maldives. Indian Ocean
Donkey and horse on the Norfolk Broads with windmill at dawn, Norfolk, UK, October 2008
Brown bear Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August
Bubble coral shrimp (Vir philippinensis) on Bubble coral (Plerogyra sp). Anilao, Philippines
Horse mussel (Modiolus modiolus) bed with Brittlestars (Ophiothrix fragilis) and Edible crab (Cancer pagurus), Shetland, Scotland, UK, September
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
Avian feeding frenzy with a flock of White Pelicans (Pelecanus erythrorhynchos) swimming along a channel chasing school of fish
Black-and-white kitten, Loona, age 11 weeks, with black-and-white rabbit, Bandit
Little bittern (Ixobrychus minutus) watching Great warbler (Acrocephalus arundinaceus) with prey, Lake Csaj, Kiskunsagi National Park, Hungary
Black-backed jackals (Canis mesomelas) two fighting with Plains / Burchell zebra (Equus burchelli) in foreground, Kalahari Desert, Nxai Pans National Park. Botswana
White tailed sea eagle (Haliaeetus albicilla) interacting with Great egret (Ardea alba) Lake Csaj, Kiskunsagi National Park, Hungary. January
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Common lobster (Homarus gammarus) looks out from its hole amongst Dead mans fingers (Alcyonium digitatum) beneath a forest of Oarweed kelp (Laminaria hyperborea)
Julia heleconia (Dryas iulia) butterfly on head of Yacare caiman (Caiman yacare). Butterflies often land on caimans head to drink the salt from its eyes. Pantanal, Brazil
Blue shark (Prionace glauca) seen from above with pilot fish, Azores Islands, Portugal, Atlantic Ocean
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Southern Type B Killer whales (Orcinus orca) hunting Weddell seal (Leptonychotes weddelli) using coordinated wave washing technique, Antarctica
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Big-eye or Goggle-eye (Priacanthus hamrur) and Crown squirrelfish (Sargocentron diadema) on wreck colonised by corals, Red Sea, Egypt. January
Spinecheek anemonefish (Premnas biaculeatus) male with bleached anemone, Bunaken National Park, North Sulawesi, Indonesia
Hoopoe (Upupa epops) with European roller (Coracias garrulus) in the background. Pusztaszer, Hungary. May
RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Angolan giraffes (Giraffa camelopardalis angolensis) and black rhinoceros (Diceros bicornis) reflecting in waterhole at sunset, Okaukuejo, Etosha National Park, Namibia, Africa
Wild Konik horses with Starlings (Sturnus vulgaris) Odry delta reserve, Stepnica, Poland, July
Bumblebee (Bombus sp) and Honey bee (Apis mellifera) nectaring on Love-in-a-mist (Nigella damascena) flower. Anthers transferring pollen onto bee backs
Hammerkop / Hammerhead stork (Scopus umbretta) eating a toad, with an African buffalo / Cape buffalo (Syncerus caffer) in the background at night, Zimanga Private Nature Reserve, KwaZulu Natal
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp)Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos
Blue mussels (Mytilus edulis) with encrusting marine life, Sula Sgeir, Scotland. UK, July
Elegant anemone (Sagartia elegans) and Plumose anemone (Metridium senile) with fish swimming in plankton bloom in background, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Black-and-white kitten, Solo, 6 weeks, rubbing against F1b toy Goldendoodle (Golden Labrador cross Toy poodle) puppy
African hunting dogs (Lycaon pictus) pack mobbing a Spotted hyena (Crocuta crocuta). Hyenas follow the dogs and attempt to steal their kills, so the dogs harass them whenever they get a chance
Great egret (Ardea alba) at twilight, Lake Csaj, Kiskunsagi National Park, Hungary. July
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica