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Grey whale (Eschrichtius robustus) pod migrating north, aerial view. Baja California, Mexico. February
Split level of Sockeye salmon (Oncorhynchus nerka) during migration; Adams River, British Columbia, Canada. October
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Trumpeter Swan (Cygnus buccinator) on water during migration through Denali mountainous tundra area, Denali Road in Alaska, North America. September
Trumpeter Swans (Cygnus buccinator) on lake, during migration through Denali mountainous tundra area, Denali Road in Alaska, North America. September
Redstart (Phoenicurus phoenicurus) perched on a barbed wire fence during autumn migration. Norfolk, UK, September (non-ex)
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Gray whale (Eschrichtius robustus) aerial view of blowing during whale migration, Coast of California, USA, Eastern Pacific Ocean
North Pacific right whale (Eubalaena japonica) swimming near surface, aerial view, Channel Islands National Park, California, USA. February
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Alewife fish (Alosa pseudoharengus), migrating up river, Northern Maine, USA. June
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Caribou (Rangifer tarandus) herd migrating across ice, Agapa River, Taimyr Peninsula, Siberia, Russia
Migrating Salmon (Salmonidae) Lake Kuril, Kamchatka, Far East Russia, August
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
Small flock of Dunlin (Calidris alpina) with a Sanderling (Calidris alba) flying in front of crashing waves. Druridge Bay, Northumberland, England, UK, January
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Grants zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya
Bewicks swans (Cygnus columbianus) small flock in flight, Gloucestershire, UK February
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
African elephants (Loxodonta africana) large family group on migration, Amboseli National Park, Kenya
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Barn swallow (Hirundo rustica) group over the cliffs of southern England (digitally enhanced)
Two Atlantic salmon (Salmo salar) silhouetted against sky, Gaspe Peninsula, Quebec, Canada, October
