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Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
White stork (Ciconia ciconia) in tree, migrating towards Europe, Masai-Mara Game Reserve, Kenya
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Atlantic salmon (Salmo salar) leaping up waterfall to reach spawning grounds upstream. River Endrick, Loch Lomond and The Trossachs National Park, Scotland, UK. July
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Eastern White-bearded Wildebeest (Connochaetes taurinus) migrating herds, Masai Mara National Reserve, Kenya
Eastern White-bearded Wildebeest herd (Connochaetes taurinus) crossing the Mara River. Masai Mara National Reserve, Kenya
Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Common crane (Grus grus) flock in flight, Lac du Der, France, November
Common / Eurasian cranes (Grus grus) flying from roost site at sunrise, silhouetted against dawn sky, autumn migration period, Rugen-Bock-Region, Mecklenberg-Vorpommern, Germany, October 2009
Adelie penguin (Pygoscelis adeliae) walking on ice, near Davis Station, Prydz Bay, East Antarctica, November
Chum salmon (Oncorhynchus keta) migrate up a small river near Bella Bella, British Columbia, Canada. September
Alewife fish (Alosa pseudoharengus), migrating up river, Northern Maine, USA. June
Zebras (Equus burchelli) herd during the Great Migration, Masai Mara National Park, Kenya. July
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Grizzly bears (Ursus arctos) on a waterfall, waiting for leaping Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Aerial view of migrating Wildebeest (Connochaetes taurinus) on the border of Ngorongoro Conservation Area and Serengeti National Park, Tanzania, Africa
Atlantic salmon (Salmo salar) leaping up waterfall, Cairngorms National Park, Scotland, UK. October
Flock of Sanderlings (Calidris Alba) in flight with wind turbines in background, Atlantic Coast, Vendee, France. December
Male Ring ouzel (Turdus torquatus) standing on a garden lawn, Norfolk, UK. April
Common cranes (Grus grus) flock on migration at sunset, Montier en Der, Aube, France, November
Flock of Snow Geese (Chen caerulescens atlanticus / Chen caerulescens) in flight and on ground at wintering area, Bosque del Apache, New Mexico, USA, November
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Northern Serenget National Park, Tanzania (early September)
Grey whale (Eschrichtius robustus) pod migrating north, aerial view. Baja California, Mexico. February
Split level of Sockeye salmon (Oncorhynchus nerka) during migration; Adams River, British Columbia, Canada. October
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Trumpeter Swan (Cygnus buccinator) on water during migration through Denali mountainous tundra area, Denali Road in Alaska, North America. September
Trumpeter Swans (Cygnus buccinator) on lake, during migration through Denali mountainous tundra area, Denali Road in Alaska, North America. September
Redstart (Phoenicurus phoenicurus) perched on a barbed wire fence during autumn migration. Norfolk, UK, September (non-ex)
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Gray whale (Eschrichtius robustus) aerial view of blowing during whale migration, Coast of California, USA, Eastern Pacific Ocean
North Pacific right whale (Eubalaena japonica) swimming near surface, aerial view, Channel Islands National Park, California, USA. February
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Caribou (Rangifer tarandus) herd migrating across ice, Agapa River, Taimyr Peninsula, Siberia, Russia
Migrating Salmon (Salmonidae) Lake Kuril, Kamchatka, Far East Russia, August
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Portrait of Male Sockeye salmon (Oncorhynchus nerka) showing teeth. Adams River, British Columbia, Canada, October
Wildebeest (Connchaetes taurinus) running down bank of the Mara River during migration, Masai Mara, Kenya
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
Atlantic salmon (Salmo salar) migrating for spawning in river, Gaspe Peninsula, Quebec, Canada, October
Low angle view of Demoiselle crane (Anthropoides virgo) flying and feeding during winter migration. Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) low angle view of birds flying and landing near the chugga ghar (bird feeding enclosure ) in Khichan Village, durin their annual migration
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Small flock of Dunlin (Calidris alpina) with a Sanderling (Calidris alba) flying in front of crashing waves. Druridge Bay, Northumberland, England, UK, January
Snow goose (Anser caerulescens) two during migration with dark clouds against the sun, Quebec, Canada, October
Atlantic salmon (Salmo salar) migrating upstream to spawn, Umelven, Sweden, July 2009
Barnacle Geese (Branta leucopsis) in flight at sunset, Caerlaverock WWT, Scotland, Solway, UK, January
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Bewicks swans (Cygnus columbianus) small flock in flight, Gloucestershire, UK February
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Massing herds of White bearded wildebeest (Connochaetes taurinus albojubatus) on migration, Serengeti National Park, Tanzania
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Atlantic salmon (Salmo salar) leaping upstream in river, Scotland, UK
African elephant (Loxodonta africana) herd walking in line, female matriach at the front, Amboseli GR, Kenya, East Africa
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Grants zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
African elephants (Loxodonta africana) large family group on migration, Amboseli National Park, Kenya
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Barn swallow (Hirundo rustica) group over the cliffs of southern England (digitally enhanced)
Two Atlantic salmon (Salmo salar) silhouetted against sky, Gaspe Peninsula, Quebec, Canada, October
