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Wild Mustang horses running, Sand Wash Basin Herd Area, Colorado, USA
Greylag geese (Anser anser). Cley, Norfolk, UK, March. Digital composite
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Yellow poppywort (Meconopsis integrifolia) flowering, Qinghai-Tibet Plateau, Shiqu County, Sichuan Province, China, August
Roseate Spoonbills (Platalea ajaja) in flight, Pantanal, Mato Grosso do Sul, Brazil
Hyacinth Macaw (Anodorhynchus hyacinthinus) small group in flight, Brazil
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Magnolia Rustica rubra in flower in garden, UK, May
Lambs in meadow in spring, UK, April
Cornflowers (Centaurea cyanus) in fallow fields, near Sugano, Orvieto, Italy, June
A flock of Mealy Amazon Parrot (Amazona farinosa) eating clay. Yasuni National Park, Orellana, Ecuador
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Pasque flowers (Pulsatilla vulgaris) in rain, Lorraine, France, April. 3rd Prize in the Other Nature Topics category of Melvita Nature Images Awards 2013
Water lily (Nymphaea rudgeana) leaves viewed from below in a black water creek. Suriname, October
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Marine Iguanas (Amblyrhynchus cristatus) on the south coast of Espanola Island, Galapagos, Vulnerable species
Lemon shark (Negaprion brevirostris) in shallow water at the surface, split level. Little Bahama Bank, Bahamas. West Atlantic Ocean
Dalmatian pelican (Pelecanus crispus) group of five resting on the lake, with mountain shrouded in mist in the background. Lake Kerkini, Greece. February. Vulnerable species
RF- Pod of Atlantic spotted dolphins (Stenella frontalis) over shallow sand bank accompanied by larger Bottlenosed dolphin (Tursiops truncatus). Sandy Ridge, Little Bahama Bank, Bahamas
RF - Banded darter (Sympetrum pedemontanum) group of five resting, Netherlands, September
A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Five Grizzly bears (Ursus arctos) standing at top of waterfall fishing for Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Icelandic horses in meadow of flowering buttercups (Ranunculus sp) southwest Iceland. June
Three Sri Lankan elephants (Elephas maximus maximus) with two calves walking along forest road, Yala National Park, Southern and Uva Provinces, Sri Lanka
Common ostriches (Struthio camelus), male and female, standing at waterhole in front of Grant's zebras (Equus quagga boehmi). Tsavo East National Park, Kenya
Four male Silver-washed fritillary butterflly (Argynnis paphia) aggregating on wildflower, Finland. July
Group of Grey partridge (Perdix perdix) huddled for warmth in snowy field. Near Nijmegen, the Netherlands. February
Giant coreopsis (coreopsis gigantea) flower, Zapato Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Lusitano horses, five young stallions playing between trees in pasture, silhouetted at dawn. Portugal. April
RF - Ring-tailed lemurs (Lemur catta) back lit and sun bathing at first light. Berenty Private Reserve, southern Madagascar. (This image may be licensed either as rights managed or royalty free.)
Arctic fox cubs (Alopex lagopus) Hornvik, Westfjords, Iceland. July
Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Incarvillea (Incarvillea compacta) with raindrops on petals. Near Zhedou Shan Pass, Sichuan Province, China
Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Pronghorn antelope (Antilocapra americana) females, Saskatchewan, Canada
Red-crowned Cranes (Grus japonicus) flying at twilight, Hokkaido, Japan, February
Brown Hare (Lepus europaeus), Bayern, Germany. April
Great egret (Ardea alba) group of five at waters edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Bearded tit (Panurus biarmicus), five perched on Reed (Phragmites australis). Danube Delta, Romania. May
King penguin (Aptenodytes patagonicus) group on rocks, jumping into South Atlantic. St Andrews Bay, South Georgia. October
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December