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Queen conch (Lobatus gigas) laying eggs in the Exuma Cays Land and Sea Park, Exuma, Bahamas
Sand dunes, marram grass (Ammophila arenaria) and beach at sunrise, Luskentyre, Isle of Harris, Scotland, UK. October 2018
RF - The Cobb, breakwater, Lyne Regis, Dorset, UK. January 2016
Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May
Sea otter (Enhydra lutris) floating on back amongst kelp, yawning, Barkley Sound, Vancouver Island, British Columbia, Canada
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Clownfish (Amphiprion sp) in anemone home, Philippines
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica
Sugar kelp (Saccharina latissima) and Sea anemones (Actiniaria) on rock near iceberg. Tasiilaq, East Greenland. April
Sugar kelp (Saccharina latissima) near iceberg, view upwards to gap in sea ice. Tasiilaq, East Greenland. April
Munks pygmy devil ray / Munks mobula (Mobula munkiana) large school from the air, Baja California, Mexico. Large group gathered possibly for mating purposes
Sea otter (Enhydra lutris) group of four resting among sea ice, Alaska, USA, June
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Harbour porpoises (Phocoena phocoena) - rare picture of small group Bay of Fundy, New Brunswick, Canada
Humpback whales (Megaptera novaeangliae) aerial. Baja California, Mexico
Blue whales (Balaenoptera musculus) surfacing, aerial shot, Baja California, Mexico
Brydes whale (Balaenoptera edeni) aerial view, Baja California, Mexico
Humpback Whale (Megaptera novaeangliae) blowing or spouting and fluking, Southeast Alaska, USA, August
Harbour porpoise (Phocoena phocoena) surfacing Bay of Fundy, New Brunswick, Canada
Spinetail devil rays (Mobula mobular) aerial Baja California, Mexico
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Australian sea lion (Neophoca cinerea), two females greeting each other on beach. Seal Bay Conservation Park, Kangaroo Island, South Australia
Lemon shark pup (Negaprion brevirostris) in mangrove forest which acts as a nursery for juveniles of this species. Eleuthera, Bahamas
Lemon shark (Negaprion brevirostris) adult resting and allowing a small cleaner wrasse feed off parasites. Grand Bahama Island, Bahamas
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Lava flow from Kilauea Volcano flowing into the Pacific Ocean, Kalapana Coast, Big Island, Hawaii. August 2016
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Whale shark (Rhincodon typus) viewed from above, Cenderawasih Bay, West Papua. Indonesia
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Storm waves off the coast of Praia do Norte, Nazare, Portugal. January
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Norfolk Island pine trees (Araucaria heterophylla) near lagoon with Mount Lidgbird and Mount Gower in the background, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site
Soft corals (Dendronephthya sp) in coral reef, West Papua, New Guinea
Commensal emperor shrimp (Periclimenes imperator) hitching a ride on a Nudibranch (Risbecia tryoni). Lembeh Strait, North Sulawesi, Indonesia
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean