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Spotted Wobbegong Shark (Orectolobus maculatus) lying in seaweed. Manly, Sydney, New South Wales, Australia. Pacific Ocean. November
Polar bear (Ursus maritimus) tagged female with wet matted fur walks over newly forming pack ice, Beaufort Sea, offshore from the 1002 area of the Arctic National Wildlife Refuge, Alaska
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA
Atlantic halibut (Hippoglossus hippoglossus) portrait, Saltstraumen, Bod, Norway, October 2008
Oldshoremore Beach and dunes in evening light, Kinlochbervie, Sutherland, Scotland, UK, April 2014
Warebeth Beach at dawn with view to Hoy, Orkney, Scotland, UK, November 2014
Rough seas during Storm Ruth, Ile d Ouessant, Armorique Regional Park. Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Polar bear (Ursus maritimus) juvenile shaking off snow after rolling around on the pack ice, arctic coast, 1002 area of the Arctic National Wildlife Refuge, Alaska, Beaufort Sea
Portrait of Polar bear (Ursus maritimus) sow standing with her cub on the snow in late winter, Arctic coast of Alaska
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Common dolphin (Delphinus delphis) jumping, Pico, Azores, Portugal, June 2009
Dusky dolphin {Lagenorhynchus obscurus} leaping at surface, Kaikoura, South Island, New Zealand
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands
Green turtle (Chelonia mydas) feeding on on Turtlegrass (Thalassia testudinum) seagrass bed. The Bahamas
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Fireworks at Peterhead harbour, Scotland, UK, July 2016
Humpback whale (Megaptera novaeangliae) two breaching at the same time together, Puerto Lopez, Santa Elena Peninsula, Manabi Province, Ecuador, July, Puerto Lopez, Santa Elena Peninsula
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Green turtle (Chelonia mydas) with rays of sunlight, black and white image, Akumal, Caribbean Sea, Mexico, July. Second place in the Visions of our Nature 2018
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Aerial shot of the Laguna Sian Ka an, freshwater lagoon, at edge of the sea, close to Boca Paila, Quintana Roo, Yucatan Peninsula, Mexico. Photographed for The Freshwater Project February 2014
Illuminated netdevil (Linophyne arboritera), male anglerfish attached to much larger female, captive from deep sea
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Bulb tentacle sea anemone (Entacmaea quadricolor) close up detail, Maldives, Indian Ocean
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
Sea Otter floating at sea surface holding pup. Alaska (Enhydra lutris) USA
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Squid (Abraliopsis atlantica) deep sea species from Atlantic Ocean off Cape Verde. Captive
Hydromedusa / Trachymedusa (Crossota sp) showing tentacles, from approx 2700m, mid Atlantic ridge, June 2010
Deepsea cirrate octopod {Stauroteuthis syrtensis} 830m, Gulf of Maine, Atlantic
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Aerial view of tidal channels in marshland, with tree like appearance. Mockhorn Island State Wildlife Management Area, Virginia, USA. May 2019
Tabular iceberg floating in Weddell Sea, iceberg broken away from Larson C ice shelf. Adelie penguin (Pygoscelis adeliae) colony dwarfed on sea ice below. Near Danger Islands, Antarctica
Pair of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea