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Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Gannet (Morus bassanus) Shetland, England, UK. July
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Short tailed albatross (Phoebastria albatrus) subadult with fish-hook and monofilament line embedded in throat. Tsubamezaki, Torishima Island, Japan. December
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Fulmar (Fulmarus glacialis) nesting amongst flowering thrift (Armeria maritima), Great Saltee Islands, County Wexford, Republic of Ireland. May
Northern gannet (Morus bassanus), Feeding frenzy, Noss NNR, Shetland, Scotland, UK
Emperor penguin (Aptenodytes forsteri), tobogganing, Snow Hill Island, Antarctic Peninsula
Atlantic puffic, (Fratercula arctica) portrait. Machias Seal Island, Bay of Fundy, New Brunswick, Canada, August
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
RF - Gannet (Morus bassanus) close-up portrait with nesting material in beak, , Saltee Islands, Ireland. June
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway
Westland petrel (Procellaria westlandica) in flight over sea, showing the upperwing. Kaikoura, South Island, New Zealand. April. Vulnerable species
Little blue / fairy penguin (Eudyptula minor) coming ashore to breeding area after dusk, Phillip Island Nature Park, Victoria, Australia
Tufted puffin (Fratercula cirrhata) in breeding plumage. St. Paul Island, Alaska. July
Horned puffin (Fratercula corniculata) close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July
Gannet (Morus bassanus) adult turning in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Wilsons Storm Petrel (Oceanites oceanicus) feeding whilst walking on water Atlantic ocean, Madeira Portugal, August
Gannet (Morus bassanus) adult wing stretching and looking up, Shetland Islands, Scotland, UK, September
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Atlantic puffin (Fratercula arctica) in flight, Isle of Lunga, Treshnish Isles, Scotland, June
Gannet (Sula bassana) in water, Shetland, UK. July
Northern gannets (Morus bassanus) diving for fish. Bempton, Yorkshire, UK. August
Snow petrel (Pagodroma nivea) flying, Antarctica, November
Humboldt penguin (Spheniscus humboldti) group gathering at landing site. Tilgo Island, La Serena, Chile. Vulnerable species
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Yellow-eyed Penguin (Megadyptes antipodes) walking to nest hidden inland in dense vegetation. Sandy Bay, Enderby Island, Auckland Group, New Zealand Sub Antarctic Islands