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Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, July
Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Gannet (Morus bassanus) Shetland, England, UK. July
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Emperor penguin (Aptenodytes forsteri) tobogganing, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Waved albatross (Phoebastria irrorata) head portrait, Espanola Island, Galapagos, South America
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Red-footed booby (Sula sula), brown morph, Genovesa Island, Galapagos Islands
Campbell albatross (Thalassarche impavida), flying off Campbell Island (seen in background), January
Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
RF - Wandering albatross (Diomedea exulans) flying over the rugged coast of South Georgia. South Atlantic
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
White capped albatross (Thalassarche steadi) in flight at sea. Auckland Islands (Subantarctic), New Zealand, February
Black-browed albatross (Thalassarche melanophris) in flight over sea at sunrise. South Atlantic Ocean between The Falklands and South Georgia. November
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Short tailed albatross (Phoebastria albatrus) subadult with fish-hook and monofilament line embedded in throat. Tsubamezaki, Torishima Island, Japan. December
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Emperor penguin (Aptenodytes forsteri) colony, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Red Footed Booby (Sula sula) couple, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Laysan Albatross (Phoebastria immutabilis) in nest, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Puffin (Fratercula arctica) among sea thrift, Great Saltee Island, County Wexford, Ireland
Fulmar (Fulmarus glacialis) nesting amongst flowering thrift (Armeria maritima), Great Saltee Islands, County Wexford, Republic of Ireland. May
Northern gannet (Morus bassanus), Feeding frenzy, Noss NNR, Shetland, Scotland, UK
Emperor penguin (Aptenodytes forsteri), tobogganing, Snow Hill Island, Antarctic Peninsula
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
Atlantic puffin (Fratercula arctica) amongst flowering Sea campion (Silene uniflora), Skomer Island, Pembrokeshire, Wales, UK. May
Emperor penguin (Aptenodytes forsteri) close up, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
RF - Atlantic puffin (Fratercula arctica) among sea campion (Silene maritima), Skomer Island, Pembrokeshire, UK. May
White-vented storm petrel (Oceanites gracilis) feeding at sea surface, in flight. Galapagos Islands, Ecuador
Waved albatross (Phoebastria irrorata) chick, aged two weeks, begging next to parent on nest. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
