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Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Maned wolf (Chrysocyon brachyurus) searching for food, Piaui, Cerrado, Brazil, South America
Tibetan Wild Ass / Kiang, (Equus kiang) running, ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India
Portrait of a common hamster (Cricetus cricetus) Alsace, France, April, captive
Chamois (Rupicapra rupicapra) running down ridge, in early morning light. Lausons Valley, Gran Paradiso National Park, Italy, September
Blue Dutch rabbit with kitten with matching colouration
Tibetan antelope (Pantholops hodgsonii) male, Kekexili, Qinghai, Tibetan Plateau, China, November
Domestic dog, Dogo Argentino, three puppies playing with stick, France
European river otter (Lutra lutra) sleeping on rocky shore, Shetland, Scotland, UK, July
Harvest mouse (Micromys minutus) on Bramble (Rubus) plant, Devon, England, UK, May. Captive
North-western wolf (Canis lupus occidentalis) portrait, captive occurs in northwestern USA and Canada
Arctic wolf (Canis lupus arctos) walking on snow, captive
Grey wolf (Canis lupus), two animals in winter, captive, USA
Tibetan wolf (Canis lupus) in snow, Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Grey wolf (Canis lupus) running in snow, captive
Vancouver Island Grey wolf (Canis lupus crassodon) Vancouver Island, British Columbia, Canada
RF- Head portrait of a European grey wolf (Canis lupus), with a second in the background, captive, Norway, February
Humpback Whale (Megaptera novaeangliae) tail slapping, tail lob, lobtailing. Vancouver Island, British Columbia, Canada, July
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa
Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya
Alpaca Herd on Norfolk Farm, winter
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Polar bear (Ursus maritimus) portrait, female. Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Red deer (Cervus elaphus) stag close up in the snow, Scotland, March
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Lion(Panthera leo) cub playing with its mothers tail, Masai-Mara Game Reserve, Kenya. Vulnerable species
Spectacled or Andean bear (Tremarctos ornatus) Maquipucuna, Pichincha, Ecuador
Portrait of Giant panda cub (Ailuropoda melanoleuca) captive. Yuan Meng, first Giant panda ever born in France, now aged 8 months, Beauval Zoo, France
Black bear (Ursus americanus), preparing for hibernation. Maine, USA
North American porcupine (Erethizon dorsatum), feeding on a young spruce tree. Vermont, USA. (Habituated rescued individual returned to the wild)
Proboscis monkey (Nasalis larvatus) female feeding, Kinabatangan River, Sabah, Borneo
Mountain tree shrew (Tupaia montana) feeding on nectar secreted by the endemic Pitcher Plant (Nepenthes kinabaluensis) Montane forests (at 2200m-3000m), slopes of Mt Kinabalu
Black-backed jackal (Canis mesomelas) three cubs at the den, Masai-Mara Game Reserve, Kenya
Bottlenose dolphin (Tursiops truncatus) at surface, Moray Firth, Highlands, Scotland. May
RF - Arctic fox (Vulpes lagopus) male, blue colour morph. Hornstrandir Nature Reserve, Iceland. March
Skull of male Urial sheep (Ovis vignei) on snow covered slope. Himalayas near Ulley, Ladakh, India
Wolf (Canis lupus) male on rocky snow covered slopes. Ulley Valley in the Himalayas, Ladakh, India
Two Jaguars (Panthera onca) snarling at each other on the riverbank, Pantanal, Brazil. September
Harvest mouse (Micromys minutus) climbing on frosty seedhead, Hertfordshire, England, UK, January, Controlled conditions
Harvest mice (Micromys minutus) on grass stems, Devon, UK. July 2016. Captive
Hedgehog (Erinaceus europaeus). Dorset, UK, July. Captive
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
Red squirrel (Sciurus vulgaris) stood on log in snow, Cairngorms National Park, Scotland, UK. December
Red Squirrel (Sciurus vulgaris) in summer coat running across fallen log in heather, Scotland, UK. September
