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Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Siberian flying squirrel (Pteromys volans orii) feeding on Sakhalin fir tree. Hokkaido, Japan. March
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Polar bear (Ursus maritimus) standing on ice. Svalbard, Norway, July
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Lion (Panthera leo) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa
RF - African elephants (Loxodonta africana) in water, trunks touching, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Siberian weasel (Mustela sibirica) standing in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Giant river otter (Pteronura brasiliensis) resting on fallen tree trunk above Cuiaba River. Pantanal, Mato Grosso, Brazil
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Melanistic leopard / Black panther (Panthera pardus fusca) resting in tree. Nagarhole National Park, India. Photo© Phillip Ross/Felis Images
Fishing cat (Prionailurus viverrinus) walking with fish in mouth. Andhra Pradesh, India. Camera trap image. Photo© Anjani Kumar/Felis Images
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Golden langur (Trachypithecus geei) sitting in tree, Umanada Temple Island, Assam, India
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Western lowland gorilla (Gorilla gorilla gorilla) male aged 12 years standing up at edge of Mpassa river. Captive bred in UK and released into wild along with his father in 2013
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with one wing spread, Yarra Bend Park, Kew, Victoria, Australia.ÔÇ¿
Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June
Aardvark (Orycteropus afer) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. Medium repro only
Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka
Giant river otter (Pteronura brasiliensis) eating a catfish. Yasuni National Park, Orellana, Ecuador
Jungle cat (Felis chaus) at night, Kanha National Park and Tiger Reserve, Madhya Pradesh, India
RF - European ground squirrel / Souslik (Spermophilus citellus) in grass with daisies, Gerasdorf, Austria. April
RF - European hamster (Cricetus cricetus) Austria
RF - European hamster (Cricetus cricetus) with cheek pouches full, Austria
Humpback whale (Megaptera novaeangliae) mother with young calf in tropical sheltered coastal waters Vava u, Tonga
Galapagos sea lion (Zalophus wollebaeki) juvenile on beach, Galapagos
A female American black bear (Ursus americanus) and her young cub, Nova Scotia, Canada. July