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RF - Lion (Panthera leo) male on savanna with dramatic storm clouds, thundery sky and light rays in the background. Ngorongoro Crater, Tanzania. Composite image
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka climbing a tree, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Elephant (Loxodonta africana) herd walking in a line, Etosha National Park, Namibia
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Cheviot sheep grazing eroding machair at front of sand dunes, North Uist, Scotland, UK, June
Adult Mountain gorilla (Gorilla beringei beringei) holding baby, Hirwa group, Volcanoes National Park, Rwanda
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
European Beaver (Castor fiber) eating blackberries by standing on its hind legs and using its tail as additional support. Part of captive breeding population prior to release into the Welsh landscape
Humpback whale (Megaptera novaeangliae) two breaching at the same time together, Puerto Lopez, Santa Elena Peninsula, Manabi Province, Ecuador, July, Puerto Lopez, Santa Elena Peninsula
Lagotto Romagnolo dog
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Meerkats (Suricata suricatta) with young, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Snow leopard (Panthera uncia) in snow, captive, USA
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Guanaco (Lama guanicoe), Punta Norte, Peninsula Valdes, Argentina
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Red panda (Ailurus fulgens) captive, occurs in China
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
Grey wolf (Canis lupus) adult greeted by cub, captive, USA
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Impala with Oxpeckers. Kruger National Park, South Africa
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
