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Moose (Alces alces) female with twin calves, Baxter State Park, Maine, USA, June
Lion (Panthera leo) close up of teeth while its snarling, captive
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
English springer spaniel in Rhododendron. Haddam, Middlesex, Connecticut, USA
Weimaraner in front of Rhododendron flowers, Haddam, Connecticut, USA. May
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
Red fox (Vulpes vulpes) visiting woodland stream to drink at night. Camera trap image. Near Bath, England, UK. June
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Grey whale (Eschrichtius robustus) breaching, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Purebred Berkshire piglet amongst autumn foliage, Smithfield, Rhode Island, USA
Wild boar (Sus scrofa) Vosges, France, July
Warthog (Phacochoerus aethiopicus) female with piglets, Okavango Delta, Botswana
Collared peccary (Tayassu tajacu), adult drinking, South Texas, USA. May
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
Wild boar (Sus scrofa) camera trap image in the Jura mountains, Switzerland, September
Purebred Berkshire piglet in oak leaves, Smithfield, Rhode Island, USA
Purebred Berkshire piglet in autumn, Smithfield, Rhode Island, USA
Collared peccary (Pecari tajacu) Laredo Borderlands, Texas, USA. April
Purebred Berkshire piglet in spring grass, dandelions and garden flowers, Smithfield, Rhode Island, USA
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Warthog (Phacochoerus africanus) mother with baby on back sleeping, Masai Mara, Kenya
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Wild boar (Sus scrofa) piglets (known as humbugs ) in woodland clearing. Forest of Dean, Gloucestershire, UK. May
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
RF - Red Dachshund puppy lying down
Black-and-tan Dachshund puppy walking
Cavapoo dog, Monty, 10 months, running
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May
Gurskys spectral tarsier (Tarsius spectrumgurskyae) North Sulawesi
Horsfields tarsier / Western tarsier ( Tarsius bancanus ssp. saltator) Belitung Island, Sumatra, Indonesia. Endemic
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
African wild dog (Lycaon pictus) portrait, Mana Pools National Park, Zimbabwe
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Red fox (Vulpes vulpes) adult with train in background, Kent, UK. March. Highly commended in the Nature Photographer of the Year Awards 2018. Taken with remote camera
White rhinoceros (Ceratotherium simum) five calves, orphaned from poaching feeding from a trough at dusk at the Rhino Revolution orphanage near Hoedspruit, South Africa. May 2017
Scarred face of a white rhinoceros (Ceratotherum simum) that survived an attack by poachers who illegally tranquillised her, two other rhinoceroses died from their injures
Aerial view of a herd of wild White rhinoceros (Ceratotherium simum) running free on Chiefs Island, Okavango Delta, Botswana
Young White rhinoceros (Ceratotherium simum) in a reinforced steel boma, blindfolded and partially drugged after a long journey from South Africa, before being released into the wild in Botswana
Mountain hare (Lepus timidus) in extreme winds with spin drifts of snow, Cairngorms National Park, Scotland, UK, February 2018
Pallass cat (Otocolobus manul) Tibetan Plateau, Qinghai, China
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Black spider monkey (Ateles chamek) calling, captive, Peru
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Black spider monkey (Ateles chamek) portrait, captive, Peru
Golden-mantled tamarin (Saguinus tripartitus) captive