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RF- Florida manatee (Trichechus manatus latirostrus) portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Ceiba / Kapok tree (Ceiba trichistandra) low angle view, Macara, Loja, Ecuador
Steppe eagle (Aquila nipalensis) in flight, Oman, November
Velvet shank / Winter fungus (Flammulina velutipes), growing on dead tree. Peak District National Park, UK. December. Focus stacked image
Hooded nudibranchs (Melibe leonina) filter feeding beneath bull kelp. Hussar Point, Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
European white waterlilies (Nymphaea alba) viewed from underwater in a tributary of Danube Delta, Romania, June
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Suwanee freshwater turtle (Pseudemys concinna suwanniensis) swimming beneath trees in a freshwater spring. Devil Spring, Ginnie Springs, High Springs, Florida, USA
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Bamboo forest, Shikoku Mura open air museum, Takamatsu, Shikoku Island, Japan, May
Dalmatian pelican (Pelecanus crispus) low angle perspective of open bill, Lake Kerkini, Greece, February. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Beech woodland in dense fog. Derbyshire, UK, November
A silhouette of a California sealion pup (Zalophus californianus) next to cliffs. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
Green turtle (Chelonia mydas) with rays of sunlight, Akumal, Caribbean Sea, Mexico, January. Endangered species
Wood Anemones (Anemone nemorosa) carpeting a woodland floor, low angle fish eye view, Peak District National Park, Derbyshire, UK, May
Canopy of a Beech woodland (Fagus sylvatica) in autumn through a circular fisheye lens. Plitvice Lakes National Park, Croatia. November
Low angle view of trees in the Hoh Rainforest, Hall of Mosses Trail, Olympic National Park, Jefferson County, Washington, USA, June
Rothschild's giraffes (Giraffa camelopardalis rothschildi), feeding on leaves, Woburn Safari Park, UK, June, captive
RF - View up Quiver tree (Aloidendron dichotomum), Western Cape, South Africa
Elephant (Loxodonta africana) at waterhole drinking at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Two Atlantic spotted dolphins (Stenella frontalis) over a shallow sand bank. On the left is a juvenile which does not have the characteristic spots of the adults
Tiger shark (Galeocerdo cuvier) swimming over sandy seabed, Tiger Beach. Bahamas, Atlantic Ocean
Maned wolf (Chrysocyon brachyurus) setting out to hunt in grassland at dusk. Serra da Canastra National Park, Brazil
Fox (Vulpes vulpes) drinking water from a sauna pool in a garden; low angle view looking up through water, Verteskozma, Hungary
Giant sequoia (Sequoiadendron giganteum) tree in forest at night, view towards canopy and sky. Sequoia and Kings Canyon National Parks, California, USA. October 2013
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Duckbill ray (Pteromylaeus bovinus) Tenerife, Canary Islands
A vibrant Caribbean coral reef with two Reef sharks (Carcharhinus perezi) and Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Aurora Borealis above silhouetted trees, northern Finland, March
Underwater view of Lake Neuchatel, close to Champ-Pittet, Canton of Vaud, Switzerland, April. Photographed for The Freshwater Project
Milky Way over an English oak tree (Quercus robur), at night, Brecon Beacons National Park International Dark Sky Preserve, Wales UK, December
Milos Viper (Macrovipera schweizeri) sub adult luring potential prey to within striking range with tail. Milos Island, Greece, July. Controlled conditions
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt
Pesquets parrot (Psittrichas fulgidus) perched on branch. Beauval Zoo Parc, France. Captive
Figwort (Scrophularia grandiflora) against sky. Globose flower with protruding upper lip that functions like umbrella to protect stamens. Cultivated in garden
Moso bamboo (Phyllostachys edulis), view upwards into canopy, Shunan Zhuhai National Park, Sichuan Province, China
Central bearded dragon (Pogona vitticeps) basking in mallee / heathland habitat on the Murrayville Track in the Big Desert Wilderness of north-western Victoria, Australia
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Crown daisy (Glebionis coronarium) flowers against sky, low angle view. Cyprus. April
Bengal tiger (Panthera tigris tigris) dominant male (T29) and resident female (T27) cooling off in a watering hole Kanha National Park, Central India. Camera trap image
Water lilies at the surface in Cenote Nicte-Ha, Tulum, Quintana Roo, Yucatan Peninsula, Mexico
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Roe deer (Capreolus capreolus) young male walking through snow. Close-up, portrait. Norway, February
Crocodile monitor (Varanus salvadorii) portrait, captive, occurs in New Guinea