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Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
Silver Birch (Betula pendula) trees in autumn colour, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Tree ferns (Cyatheaceae) in Kells Bay Gardens, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
A Florida manatee (Trichechus manatus latirostrus) head portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Watermilfoil (Myriophyllum spicatum) with some stems supporting clusters of Zebra mussel (Dreissena polymorpha) Lake Neuchatel, Switzerland
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Green sea turtle (Chelonia mydas) swimming near surface in halo of light. The Bahamas
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
Beech (Fagus sylvatica) forest, view into canopy from below. Peerdsbos, Brasschaat, Belgium. April 2019
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Honey Fungus (Armillaria mellea) growing on mature Ash tree (Fraxinus excelsior). Peak District National Park, Derbyshire, UK. September
Bengal tiger (Panthera tigris tigirs) tigress, stopping due to sound from camera while patrolling, Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigris) (T27) walking on bund / dam wall trail. Kanha National Park, Central India. Camera trap image
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October
RF - Dandelion (Taraxacum officinale) seeds blowing in the wind, England, UK. May
Sugar kelp (Saccharina latissima) near iceberg, view upwards to gap in sea ice. Tasiilaq, East Greenland. April
Himalayan marmot (Marmota himalayana). Chantang Wlidlife Sanctuary. Ladakh, India
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Spanish cedar (Cedrela odorata) tree, Manu National Park, Peru
Close-up of the gills of a group of small mushrooms (unidentified) north Cornwall, UK. November. Highly commended in the Wild Woods Category of the BWPA
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
Northern pitcher plant (Sarracenia purpurea) photographed on Borgles Island, Nova Scotia, Canada, September
Red oak (Quercus rubra) low angle view of trees, New Brunswick, Canada, May
Royal python (Python regius) Togo. Controlled conditions
RF- Florida manatee (Trichechus manatus latirostrus) portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Ceiba / Kapok tree (Ceiba trichistandra) low angle view, Macara, Loja, Ecuador
Steppe eagle (Aquila nipalensis) in flight, Oman, November