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Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Japanese dwarf flying squirrel (Pteromys volans orii) gliding through forest canopy during light snowfall. Hokkaido, Japan. February
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Tree ferns (Cyatheaceae) in Kells Bay Gardens, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Silver Birch (Betula pendula) trees in autumn colour, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Golden eagle (Aquila chrysaetos) sub-adult flying, Strathdearn, Inverness-shire, Scotland, UK, August
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
A Florida manatee (Trichechus manatus latirostrus) head portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Watermilfoil (Myriophyllum spicatum) with some stems supporting clusters of Zebra mussel (Dreissena polymorpha) Lake Neuchatel, Switzerland
Giant sequoia (Sequoiadendron giganteum) tree in forest at night, view towards canopy and sky. Sequoia and Kings Canyon National Parks, California, USA. October 2013
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Green sea turtle (Chelonia mydas) swimming near surface in halo of light. The Bahamas
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
Beech (Fagus sylvatica) forest, view into canopy from below. Peerdsbos, Brasschaat, Belgium. April 2019
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Central bearded dragon (Pogona vitticeps) basking in mallee / heathland habitat on the Murrayville Track in the Big Desert Wilderness of north-western Victoria, Australia
Honey Fungus (Armillaria mellea) growing on mature Ash tree (Fraxinus excelsior). Peak District National Park, Derbyshire, UK. September
Bengal tiger (Panthera tigris tigris) dominant male (T29) and resident female (T27) cooling off in a watering hole Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigirs) tigress, stopping due to sound from camera while patrolling, Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigris) (T27) walking on bund / dam wall trail. Kanha National Park, Central India. Camera trap image
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October
RF - Dandelion (Taraxacum officinale) seeds blowing in the wind, England, UK. May
Sugar kelp (Saccharina latissima) near iceberg, view upwards to gap in sea ice. Tasiilaq, East Greenland. April
Himalayan marmot (Marmota himalayana). Chantang Wlidlife Sanctuary. Ladakh, India
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Caracal (Caracal caracal) male, standing on rock against blue sky, Spain. Captive, occurs in Africa and Asia
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Fosa (Cryptoprocta ferox) pair, mating in forest canopy, Zombitse-Vohibasia National Park, central south western Madagascar
Tiger shark (Galeocerdo cuvier) swimming over sandy seabed, Tiger Beach. Bahamas, Atlantic Ocean
Grey-headed flying-fox bat (Pteropus poliocephalus) hanging from branch looking down with wing extended, Yarra Bend Park, Victoria, Australia. Cropped
Old male Espanola giant Galapagos tortoise (Chelonoidis hoodensis) stretching its neck in clearing, Las Tunas, Espanola Island, Galapagos Islands, Ecuador
Maned wolf (Chrysocyon brachyurus) setting out to hunt in grassland at dusk. Serra da Canastra National Park, Brazil
Fox (Vulpes vulpes) drinking water from a sauna pool in a garden; low angle view looking up through water, Verteskozma, Hungary
Female Red fox (Vulpes vulpes) foraging inside a rotting tree trunk, Verteskozma, Hungary. Hungary Nature Photographer of the Year Competition 2021 Highly Commended - Mammals Behaviour category
Spider crab (Hyas sp. ) scaling the stipe of Cuvie kelp (Laminaria hyperborea)Spider crab (Hyas sp.) scaling the stipe of Cuvie kelp (Laminaria hyperborea) up into the canopy of the kelp forest. Loch Carron, Highlands, Scotland, UK
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
A lesser horseshoe bat (Rhinolophus hipposideros) flying over grassland habitat at night. Devon, UK
Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Duckbill ray (Pteromylaeus bovinus) Tenerife, Canary Islands
A vibrant Caribbean coral reef with two Reef sharks (Carcharhinus perezi) and Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Aurora Borealis above silhouetted trees, northern Finland, March
Underwater view of Lake Neuchatel, close to Champ-Pittet, Canton of Vaud, Switzerland, April. Photographed for The Freshwater Project
Milky Way over an English oak tree (Quercus robur), at night, Brecon Beacons National Park International Dark Sky Preserve, Wales UK, December
Milos Viper (Macrovipera schweizeri) sub adult luring potential prey to within striking range with tail. Milos Island, Greece, July. Controlled conditions
Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt
Pesquets parrot (Psittrichas fulgidus) perched on branch. Beauval Zoo Parc, France. Captive
Figwort (Scrophularia grandiflora) against sky. Globose flower with protruding upper lip that functions like umbrella to protect stamens. Cultivated in garden
Moso bamboo (Phyllostachys edulis), view upwards into canopy, Shunan Zhuhai National Park, Sichuan Province, China
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Crown daisy (Glebionis coronarium) flowers against sky, low angle view. Cyprus. April
Water lilies at the surface in Cenote Nicte-Ha, Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Shoal of Piraputanga, (Brycon hilarii), close to the surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Roe deer (Capreolus capreolus) young male walking through snow. Close-up, portrait. Norway, February
Crocodile monitor (Varanus salvadorii) portrait, captive, occurs in New Guinea
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
European ground squirrels / Sousliks (Spermophilus citellus) in grass with daisies, Gerasdorf, Austria. April
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
King cobra (Ophiophagus hannah), low wide angle perspective Agumbe, Karnataka, Western Ghats, India. Vulnerable species
Bell glass frog (Cochranella nola) from below, photographed on a pane of glass in Manu Biosphere Reserve, Peru
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Oceanic manta ray (Mobula birostris) accompanied by three Black jacks (Caranx lugubris) silhouetted, Revillagigedo Islands, Mexico, Pacific Ocean
Ventral view of Swallowtail butterfly (Papilio machaon), male and female, in flight. The Netherlands. July
Cheetah (Acinonyx jubatus) standing on top of a rock, Spain. Captive
RF - Two Cheetahs (Acinonyx jubatus) standing and sitting on rocks against blue sky, Spain. Captive. (This image may be licensed either as rights managed or royalty free.)