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Arctic fox (Vulpes lagopus), female blue colour morph in winter coat, during snow blizzard. Baited with small amount of natural food (dried fish). Hornstrandir, Iceland. February
Mountain hare (Lepus timidus) with fur encrusted in snow, Cairngorms National Park, Scotland, UK. January
Adelie penguin (Pygoscelis adeliae) leaping out of ocean as it approaches land, Devil Island, Antarctica, Southern Ocean
Lioness (Panthera leo) lying in long grass on the savannah, portrait, Okavango Delta, Botswana, Africa
Lion (Panthera leo) cub, aged 3 months, climbing on branch and chewing on a stick, Okavango Delta, Botswana, Africa
Two Dwarf mongoose (Helogale parvula) peering out of their burrow in termite mound, Okavango Delta, Botswana, Africa
Four Common / Plains Zebras (Equus quagga) standing side by side, portrait, Okavango Delta, Botswana, Africa
Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Asian small-clawed otter (Aonyx cinerea) pups looking at camera. Captive, occurs in Asia. Zooparc Overloon, the Netherlands
Male Highland Great diving beetle (Dytiscus lapponicus) on surface of water, Darnaway Estate, Highlands, Scotland, UK. May. Focus stacked. Captive
Kerry bog pony, gelding, a rare breed, head portrait, County Kerry, Republic of Ireland. April
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
African wild dog (Lycaon pictus) alpha female suckling her 12 pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
African wild dog (Lycaon pictus) head portrait, Okavango Delta, Northern Botswana, Africa
Crab spider (Synaema globosum) on a Poppy (Papaver sp. ) flower, Sibillini, Umbria, Italy. MayCrab spider (Synaema globosum) on a Poppy (Papaver sp.) flower, Sibillini, Umbria, Italy. May
Brown-throated sloth (Bradypus variegatus) climbing on tree branch, Yasuni National Park, Orellana, Ecuador
Galapagos sea lion (Zalophus wollebaeki) diving, Wolf Island, Galapagos Islands, Pacific Ocean
Two Speckled moray eels (Gymnothorax dovii) feeding on a Sea turtle carcass on the seabed, Wolf Island, Galapagos Islands, Pacific Ocean
Peacock katydid (Pterochroza ocellata) on leaf in defensive posture, Yasuni National Park, Amazon Basin, Ecuador
Cheetah (Acinonyx jubatus) resting, wild but habituated animal, Zimanga Game Reserve, South Africa
Puget Sound King crab (Lopholithodes mandtii) extreme close-up of face, Vancouver Island, British Columbia, Canada
Patagonian / Austral pygmy owl (Glaucidium nana) perched on branch in snow, Torres del Paine National Park, Patagonia, Chile
Guanaco (Lama guanicoe) dusted in snow, head portrait, Torres del Paine National Park, Patagonia, Chile
Young male Lion (Panthera leo) resting in tree during middle of the day to escape the heat. Acacia woodland near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border, Tanzania
Young male Lion and lioness (Panthera leo) resting in tree during middle of the day to escape the heat. Acacia woodland near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border
Juvenile Puma (Puma concolor) captured on a camera trap in the Ecuadorian Choco rainforest, one of the most biodiverse and threatened rainforests on Earth. Mashpi Reserve, Pichincha, Ecuador, April
Jack snipe (Lymnocryptes minimus) resting among seaweed on rocky shoreline, Parainen Uto, Finland. September
Two male Common / Plains zebras (Equus quagga) fighting, Okavango Delta, Botswana, Africa
Giraffe (Giraffa camelopardalis angolensis) calf, aged 6 weeks, with two Yellow-billed oxpeckers (Buphagus africanus) perched on its head feeding on parasites, Okavango Delta, Botswana, Africa
Chalcid parasitoid wasp (probably Mesopolobus tibialis), 2mm in length, moving over Oak (Quercus sp)Chalcid parasitoid wasp (probably Mesopolobus tibialis), 2mm in length, moving over Oak (Quercus sp.) buds, Peak District National Park, Derbyshire, UK. April. Focus Stacked Image
Two Lionesses (Panthera leo) nuzzling surrounded by three cubs, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Giant panda (Ailuropoda melanoleuca) cub Yuandudu, aged 8 months, portrait, Beauval ZooPark, France, April, 2022. Captive
Male Jumping spider (Euophrys frontalis) close up, Derbyshire, UK. May. Focus stacked image
Green tiger beetle (Cicindela campestris) close uo portrait, Birchover, Derbyshire, UK. April. Focus stacked image
Streaked gurnard (Trigloporus lastoviza) portrait, Marine Protected area Punta Campanella, Costa Amalfitana, Italy, Tyrrhenian Sea
Common greenshank (Tringa nebularia) wading in river with wings spread, Allahein River, The Gambia
Splendid toadfish (Sanopus splendidus) emerging from its daytime hiding place at night, Cozumel, Mexico, Caribbean Sea
Buzzard (Buteo buteo) perched on a branch in autumn, Lorraine, France. November
Iberian hare (Lepus granatensis) portrait, Castile La Mancha, Spain. July
White-eyed moray (Muraena thyrsoides) close up head portrait, Bali, Indonesia, Indo-Pacific
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
Green sea turtle (Chelonia mydas) swimming over a reef, portrait, Hawaii, Pacific Ocean. Endangered
RF - Male Blackhead blenny (Emblemariopsis bahamensis) peering out from a hole in a Brain coral (Scleractinia), Grand Cayman, Cayman Islands, Caribbean Sea
RF - Grey angelfish (Pomacanthus arcuatus) portrait, Grand Cayman, Cayman Islands, Caribbean Sea. (This image may be licensed either as rights managed or royalty free.)
RF - Bluebelly blenny (Alloblennius pictus) looking out from hole in the reef, Gubal Island, Egypt, Red Sea. (This image may be licensed either as rights managed or royalty free.)
Mexican hogfish (Bodianus diplotaenia) swimming on its side, Socorro Island, Baja California, Mexico, Pacific Ocean
A tiny Revillagigedos barnacle-blenny (Acanthemblemaria mangognatha) peeking out from a crevice, Socorro Island, Baja California, Mexico, Pacific Ocean
Harp seal (Pagophilus groenlandicus) pup about four weeks old, moulting, swimming under sea ice, Magdalen Islands, Gulf of St Lawrence, Canada