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RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Bramble (Rubus plicatus) leaves and stem against white background. France, August
Panther Chameleon {Furcifer pardalis} close-up of skin, green colouration. Masoala Peninsula National Park, north east Madagascar
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Aylesbury ducks following in a line on village green, Weedon, Buckinghamshire, UK, October
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Spring crocus (Crocus vernus) flowering en masse on the Campo Imperatore, Abruzzo, Italy. April
Honeybee (Apis melifera) visiting Lavender (Lavendula angustifolia) in lavender fields, Valensole Plateau, Alpes Haute Provence, France, July
Thimble jellyfish (Linuche unguiculata) aggregation, My Paradise Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Zebras (Equus burchelli) herd during the Great Migration, Masai Mara National Park, Kenya. July
A flock of Starlings (Sturnus vulgaris) flying in front of the Blackpool Tower, England, UK, September 2010
A panoramic composite image of a massive grouping of King penguins (Aptenodytes patagonicus) seeking shelter in the lee of small hills during a blizzard
King penguins (Aptenodytes patagonicus) congregate on the beach, seeking a safe point to go into the sea, with male Southern elephant seals (Mirounga leonina) blocking their path
During an extreme heat-stress event at Melbournes Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43'C
Dulmen ponies, wild mares and foals running together at roundup. Dulmen, North Rhine-Westphalia, Germany
Slime mould (Physarum psittacinum), in mature reproductive phase. Close-up of erupting fruiting bodies (sporangia), bearing thousands of spores. Buckinghamshire, UK
Golden jellyfish (Mastigias sp. ) swarm in a marine lake, Jellyfish Lake, Eil Malk islandGolden jellyfish (Mastigias sp.) swarm in a marine lake, Jellyfish Lake, Eil Malk island, Rock Islands, Palau
Sculpin (Bero elegans) eggs. Hokkaido, Japan. May
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August
Greater rhea (Rhea americana) nest with many eggs, at edge of arable field. Patagonia, Argentina. October
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
Knot (Calidris canutus) and grey plover (Pluvialis squatarola) roost at high tide with numerous birds packed tightly together, Snettisham, Norfolk, October
Sulphur tuft mushrooms (Hypholoma fasciculare), Hampstead Heath, England, UK. September
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Avian feeding frenzy with a flock of White Pelicans (Pelecanus erythrorhynchos) swimming along a channel chasing school of fish
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Cockroaches (Periplaneta americana) on wall of Gomantong Cave. Sabah, Borneo. Malaysia
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Plankton at night, Palau, Philippine Sea
Army ants (Eciton sp. ) Costa Rica. February 2015Army ants (Eciton sp.) Costa Rica. February 2015
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Low angle view of Demoiselle crane (Anthropoides virgo) flying and feeding during winter migration. Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) low angle view of birds flying and landing near the chugga ghar (bird feeding enclosure ) in Khichan Village, durin their annual migration
Brown fur seal (Arctocephalus pusillus) hauled out on board walk in Cape Cross seal colony, Namibia
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp)Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean
Western sandpipers (Calidris mauri) roosting on barrier island tidal flats. Terrebonne Parish, Louisiana, USA. October
Lesser flamingos (Phoeniconaias minor) wading in saltwater lagoon, Salinas, Walvis Bay, Namibia
Spotted hyena (Crocuta crocuta) hunting Lesser flamingoes (Phoenicopterus minor) Lake Nakuru, Kenya