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Mallards (Anas platyrhynchos) slipping on ice on frozen lake, Wiltshire, UK, January
Asiatic lion cub (Panthera leo persica) looking up into tree, possibly at a bird, Gir Forest NP, Gujarat, India
Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
Blue-tailed damselfly {Ischnura elegans} eyes just visible behind reed stem, Cornwall, UK. June
Striped Field Mouse (Apodemus agrarius) and Corn Poppy (Papaver rhoeas). Lublin Highland, Poland, August
Cane Toad / Marine Toad / Giant Toad (Bufo marinus) adult jumping in Dogweed (Dyssodia pentachaeta) field. Laredo, Webb County, South Texas, USA, April
Two playful Bulldog puppies, 11 weeks
Four Mallard (Anas platyrhynchos) dabbling for food in icy water. The Netherlands, January
Basset hound, portrait, resting head on back of car, eyes half closed, UK
Polar bear (Ursus maritimus) young male playing / rolling around in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Head portrait of Highland cow, Scotland, with tiny frond of bracken at corner of mouth, UK
Three Ring-tailed Lemurs (Lemur catta) sun basking at dawn. Berenty Private Reserve, southern Madagascar. (digitally stitched image)
Front-view showing head and eyes of Common frog (Rana temporaria), Broxwater, Cornwall, UK
Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Polar bear (Ursus maritimus) female, resting on ice with cub sitting next to her waving and poking out tongue, Svalbard, Norway. April
Three Common toads (Bufo bufo) in amplexus, Lucerne, Switzerland. March. Cropped
Two African scops owls (Otus senegalensis) perched in tree, Katavi National Park, Tanzania
California sea lion (Zalophus californianus) close up with group playing in shallow water behind, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Lion-tailed macaque (Macaca silenus) infant, sitting among the troop with its finger in its mouth, Valparai, Tamil Nadu, India
Imbabura tree frog (Boana picturata) restin on a leaf at night, Mashpi, Pichincha, Ecuador
Male Siberian flying squirrel (Pteromys volans orii) attempting to enter nest with female inside during reproductive season but stopped mid-way then rejected. Hokkaido, Japan. March
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Black Welsh mountain sheep lamb calling. Wales UK, March
Two Florida manatees (Trichechus manatus latirostris) close to the surface in shallow water, Three Sisters Spring, Crystal River, Florida, USA. Endangered
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Adelie penguin (Pygoscelis adeliae) leaping out of ocean as it approaches land, Devil Island, Antarctica, Southern Ocean
Two Dwarf mongoose (Helogale parvula) peering out of their burrow in termite mound, Okavango Delta, Botswana, Africa
Brown-throated sloth (Bradypus variegatus) climbing on tree branch, Yasuni National Park, Orellana, Ecuador
Galapagos sea lion (Zalophus wollebaeki) diving, Wolf Island, Galapagos Islands, Pacific Ocean
Guanaco (Lama guanicoe) dusted in snow, head portrait, Torres del Paine National Park, Patagonia, Chile
White-eyed moray (Muraena thyrsoides) close up head portrait, Bali, Indonesia, Indo-Pacific
Green sea turtle (Chelonia mydas) swimming over a reef, portrait, Hawaii, Pacific Ocean. Endangered
RF - Male Blackhead blenny (Emblemariopsis bahamensis) peering out from a hole in a Brain coral (Scleractinia), Grand Cayman, Cayman Islands, Caribbean Sea
RF - Bluebelly blenny (Alloblennius pictus) looking out from hole in the reef, Gubal Island, Egypt, Red Sea. (This image may be licensed either as rights managed or royalty free.)
Mexican hogfish (Bodianus diplotaenia) swimming on its side, Socorro Island, Baja California, Mexico, Pacific Ocean
An endemic Hawaiian pom-pom crab / Boxer crab (Lybia edmondsoni) carrying a juvenile Anemone (Triactis producta) in its claws as a defense mechanism
Northern fulmar (Fulmarus glacialis) floating on surface of open ocean, Falmouth, English Channel, Cornwall, UK. September
Jaguar (Panthera onca) at night, portrait, La Papalota, Nayarit, Mexico. Camera trap image
Northern fulmar (Fulmarus glacialis) portrait, Fair Isle, Shetland, Scotland, UK. May
Spinyhead blenny (Acanthemblemaria spinosa) peeking out from a hole inhard coral, Bonaire, Lesser Antiles, Caribbean Sea
An inflated Guineafowl pufferfish (Arothron meleagris) in front of sea fans (Gorgonia sp)An inflated Guineafowl pufferfish (Arothron meleagris) in front of sea fans (Gorgonia sp.) on a reef wall, Fiji, Pacific Ocean
Spotted porcupinefish (Diodon hystrix) swimming over a reef, Hawaii, Pacific Ocean
Asiatic lion (Panthera leo persica) cub, Gir Forest National Park, Gujarat, India. May
Infant Bornean Orang-Utan (Pongo pygmaeus) clinging to its mother. Camp Leakey, Tanjung Puting NP, Kalimantan, Borneo
Western tarsier (Tarsius bancanus) clinging to tree, Danum Valley, Sabah, Borneo