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Aldabra Giant Tortoises (Aldabrachelys gigantea) resting in a pool to keep cool, Grand Terre, Natural World Heritage Site, Aldabra
Aerial view of Cape Town city with Table Mountain, South Africa, taken from helicopter, May 2011
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
Slime mould (Physarum sp) plasmodium growing across rotting wood, Osa Peninsula, Costa Rica
Spiny predatory katydid (Listroscelis sp) photographed on a white background in mobile field studio, Osa Peninsula, Costa Rica
Clearwing moth (Cosmosoma teuthras) photographed on a white background in mobile field studio, Cordillera de Talamanca mountain range, Caribbean Slopes, Costa Rica
Plankton at night, Palau, Philippine Sea
Shag (Phalacrocorax aristotelis) two heads showing above rock, Vardo, Norway March
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Okarito Brown Kiwi (Apteryx rowi) feather detail resembling mammalian hair. Okarito Forest, Westland, South Island, New Zealand
Atlas moth {Attacus atlas} on nurse plant Sri Lanka
Great white egrets (Casmerodius albus) reflected in Pulicat Lake, Tamil Nadu, India, January 2013
Warthog (Phacochoerus aethiopicus) with young, Samburu Game Reserve, Kenya
View over Llangors from Mynydd Troed, Brecon Beacons, Powys, Wales, UK. June 2014
Stonehenge at sunrise, Salisbury Plain, Wiltshire, England, UK, June 2006
Gatklettur, Arnarstapi, Snaefellsnes Peninsula, Iceland, March 2017
Southern Fjallabak, Iceland. September 2017
Belvedere at dawn, Val d Orcia, Tuscany, Italy, May 2018
Landscape of Alpe di Siusi, Dolomites, Italy, March 2016
Lago d Antorno and Cadini, Dolomites, Italy, October 2017
Lago de Predil, Julian Alps, Italy, October 2013
Sand Pyramids, Rozhen, Melnik, Bulgaria. October 2017
Riisitunturi in winter with Aurora Borealis, Kuusamo, Lapland, Finland. January 2016
Jasna Lakes in autumn, Kranjska Gora, Julian Alps, Slovenia. October 2014
Church of St. Primoz, Jamnik, Slovenia. July 2017
Church of the Assumption of St. Mary and Bled Castle, Bled Island, Julian Alps, Slovenia, January 2010
View towards Glastonbury Tor over reedbeds at dawn, Somerset, England
Cowslips (Primula veris) in bloom on Hardington Moor NNR, Somerset, England, UK, May 2013
Spinecheek anemonefish (Premnas biaculeatus) male with bleached anemone, Bunaken National Park, North Sulawesi, Indonesia
Harpy eagle (Harpia harpyja) native to South America, captive
Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Jaguar (Panthera onca) mother playing with four month old cub, native to Southern and Central America, captive
Jaguar (Panthera onca) mother grooming with four month old cub, native to Southern and Central America, captive
Wild male Jaguar in stream, Amazon Basin, Brazil S. America
Giant panda {Ailuropoda melanoleuca} Qionglai Mts, Sichaun, China
Blackback Eastern lowland gorilla {Gorilla beringei graueri} Kahuzi Biega, Congo
Juvenile Snow leopard {Panthera uncia} captive
Snow leopard (Panthera uncia) resting in snow. Captive
Polar bear (Ursus maritimus) portrait of a yearling along Bernard Spit, a barrier island, during autumn freeze up, along the eastern Arctic coast of Alaska, Beaufort Sea, September
Polar bear portrait {Ursus maritimus} Coastal plain of the Arctic National Wildlife Refuge, Alaska
Yellow-eyed penguin preening feathers, Aukland Island, New Zealand
RF - Polar Bear (Ursus maritimus), female resting, with two cubs along a barrier island outside Kaktovik, Alaska, USA. September
Dipper (Cinclus cinclus) displaying or drying wings, Kuusamo Finland March
A group of Haflinger, Pure Arab, Shagya Arab and East Bulgarian fillies and mares running in snow, Kabiuk National Stud, Shumen, Bulgaria
Two Cheetah (Acinonyx jubatus) chasing Springbok (Antidorcas marsupialis) Kgalagadi, South Africa
RF - Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway